Venerupis philippinarum
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''Lajonkairia lajonkairii'' is an edible
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of saltwater
clam Clam is a common name for several kinds of bivalve molluscs. The word is often applied only to those that are edible and live as infauna, spending most of their lives halfway buried in the sand of the seafloor or riverbeds. Clams have two shel ...
in the family
Veneridae The Veneridae or venerids, common name: Venus clams, are a very large family of minute to large, saltwater clams, marine bivalve molluscs. Over 500 living species of venerid bivalves are known, most of which are edible, and many of which are ex ...
, the Venus clams. MolluscaBase eds. (2022). MolluscaBase. Lajonkairia lajonkairii (Payraudeau, 1826). Accessed through: World Register of Marine Species at: https://marinespecies.org/aphia.php?p=taxdetails&id=140727 on 2022-02-05 Common names for the species include Manila clam, Japanese littleneck clam, Japanese cockleCohen, A. N. 2011
''Venerupis philippinarum''.
The Exotics Guide: Non-native Marine Species of the North American Pacific Coast. Center for Research on Aquatic Bioinvasions, Richmond, California, and San Francisco Estuary Institute, Oakland, California. Revised September 2011.
and Japanese carpet shell. This clam is commercially harvested, being the second most important bivalve grown in aquaculture worldwide.Cordero, D., et al
Population genetics of the Manila clam (Ruditapes philippinarum) introduced in North America and Europe.
''Nature: Scientific Reports'' 7, Article number: 39745. 3 January 2017.


Description

The shell of ''Lajonkairia lajonkairii'' is elongated, oval, and sculptured with radiating ribs.Morris, R.H., Abbott, D.P., & Haderlie, E.C. (1980). ''Intertidal Invertebrates of California''. Stanford, CA: Stanford University Press. It is generally 40 to 57 millimeters wide, with a maximum width of 79 millimeters.Fofonoff P. W., et al
''Lajonkairia lajonkairii''.
National Exotic Marine and Estuarine Species Information System (NEMESIS). Accessed 22 May 2017.
The shell is variable in color and patterning, being cream-colored to gray with concentric lines or patches. Individuals living in anoxic conditions may be black. The inside surface of the shell is often white with purple edges. The siphons are separated at the tips.Carlton, J. T. (Ed.) (2007). ''The Light and Smith Manual: Intertidal Invertebrates from Central California to Oregon. University of California Press.


Distribution

This clam is native to the coasts of the Indian,
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
and Pacific Oceans from
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...
and
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
north to China, Japan,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
and the Kuril Islands.''Ruditapes philippinarum''.
Fisheries and Aquaculture Department, FAO. 2017.
It has an extensive nonnative distribution, having been introduced accidentally and purposely as a commercially harvested edible clam. It is now permanently established in coastal ecosystems in many parts of the world. It is common along the Pacific coast of North America from
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
to
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
, where its original introduction was accidental. It can be found in
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state ...
. It was first seeded in the waters of Europe in the 1970s, and there have been multiple introductions throughout the region. It has spread naturally in Western Europe over the decades, its adaptability allowing it to thrive in many coastal habitat types. It has been planted in
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
, and French Polynesia for the purposes of aquaculture.


Habitat

This burrowing clam is most abundant in subtropical and cooler temperate areas. It can be found in shallow waters in coarse sand, mud, and gravel substrates. It lives in the littoral and sublittoral zones.Palomares, M. L. D. and D. Pauly (Eds.
''Ruditapes philippinarum''.
SeaLifeBase. Version February 2017.
It burrows no more than 10 centimeters into the substrate. It sometimes lives in
eelgrass Eelgrass is a common name for several plants and may refer to: * ''Zostera'', marine eelgrass * ''Vallisneria'', freshwater eelgrass {{Short pages monitor