Urolite is a term composed of two Greek words, ''uro-'' meaning "

urine Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excreted from the body through the urethra. Cellular ...

" and ''lithos'' meaning "

stone In geology, rock (or stone) is any naturally occurring solid mass or aggregate of minerals or mineraloid matter. It is categorized by the minerals included, its Chemical compound, chemical composition, and the way in which it is formed. Rocks ...

" and was first used to describe the fossil of a nonliquid urinary secretions produced by some groups of reptiles, in relation to

coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...

s. The first evidence of recorded liquid waste elimination attributed to a dinosaur was presented to the public in 2002, but no scientific paper had reported fossil evidence of liquid waste of

tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids ( pelycosaurs, extinct t ...

s elimination to assume that

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s urinated. In 2004, a paper by

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

Marcelo Adorna Fernandes brought a study of

trace fossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...

s that had been preserved in three aeolian

flagstone Flagstone (flag) is a generic flat stone, sometimes cut in regular rectangular or square shape and usually used for paving slabs or walkways, patios, flooring, fences and roofing. It may be used for memorials, headstones, facades and other co ...

s. These trace fossils show a pattern that could be formed by an abundant falling stream of fluid and that is different from the structures described before in other occurrences in the paleontological record. The aspect of these urolites is very similar to soil deformation caused by modern

ostrich Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There ...

urination, and certain groups of dinosaurs could have a similar urinary physiology. These urolites are the first evidence of liquid waste attributed to dinosaurs.

References

{{reflist Trace fossils