''Unescoceratops'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

leptoceratopsid Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, bu ...

ceratopsian Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s known from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

, southern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

. It contains a single species, ''Unescoceratops koppelhusae''.

Discovery

''Unescoceratops'' is known only from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

specimen

TMP TMP can refer to any of the following: Chemistry * 2,2,6,6-Tetramethylpiperidine, an organic chemistry reagent * Thymidine monophosphate, a nucleotide * Trimethoprim, an antibiotic * Trimethyl phosphate, a solvent * Trimethylolpropane, a precurso ...

95.12.6, a partial left

dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

. The fossil was collected in 1995 in the Black Coulee locality (formerly Deadhorse Coulee), near the

Writing-on-Stone Provincial Park Writing-on-Stone Provincial Park is located about southeast of Lethbridge, Alberta, Canada, or east of the community of Milk River, and straddles the Milk River itself. It is one of the largest areas of protected prairie in the Alberta park sy ...

, from the

Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...

, dating to the late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

stage of the

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

, about 76.5-75

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. The specimen was regarded as too incomplete to identify, and was shelved for several years. It was believed to be a ''Leptoceratops'' specimen at the time. A

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis done by Michael Ryan (of The Cleveland Museum of Natural History) and David Evans (of the

Royal Ontario Museum The Royal Ontario Museum (ROM) is a museum of art, world culture and natural history in Toronto, Ontario, Canada. It is one of the largest museums in North America and the largest in Canada. It attracts more than one million visitors every year ...

) found it to be among the most advanced leptoceratopsid genera.

Description

''Unescoceratops'' is thought to have been between one and two meters long and less than 91 kilograms. Its teeth were the roundest of all

s. Mallon et al. (2013) examined herbivore coexistence on the island continent of

Laramidia Laramidia was an island continent that existed during the Late Cretaceous period (99.6–66 Ma), when the Western Interior Seaway split the continent of North America in two. In the Mesozoic era, Laramidia was an island land mass separated from A ...

, during the Late Cretaceous. It was concluded that small

ornithischians Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...

like ''Unescoceratops'' were generally restricted to feeding on vegetation at, or below the height of 1 meter.

Etymology

The genus ''Unescoceratops'' was first described by Michael J. Ryan,

David C. Evans David Cannon Evans (February 24, 1924 – October 3, 1998) was the founder of the computer science department at the University of Utah and co-founder (with Ivan Sutherland) of Evans & Sutherland, a pioneering firm in computer graphics hardwar ...

Philip J. Currie Philip John Currie (born March 13, 1949) is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the ...

Caleb M. Brown Caleb (), sometimes transliterated as Kaleb ( he, כָּלֵב, ''Kalev'', ; Tiberian vocalization: Kālēḇ; Hebrew Academy: Kalev), is a figure who appears in the Hebrew Bible as a representative of the Tribe of Judah during the Israelites' ...

and

Don Brinkman Don, don or DON and variants may refer to: Places *County Donegal, Ireland, Chapman code DON *Don (river), a river in European Russia *Don River (disambiguation), several other rivers with the name *Don, Benin, a town in Benin *Don, Dang, a vill ...

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gather ...

(though it had been published earlier by Ryan and Currie in 1995 with no name) and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Unescoceratops koppelhusae''. The name means "

UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...

" (''ceratopsian'' means "one with a horned face"). The name is meant to honor UNESCO's efforts to increase understanding of natural history sites around the world. "'' Dinosaurprovincialparkaceratops'' was too long," explained Ryan. The specific epithet is named after Currie's wife,

palynologist Palynology is the "study of dust" (from grc-gre, παλύνω, palynō, "strew, sprinkle" and ''-logy'') or of "particles that are strewn". A classic palynologist analyses particulate samples collected from the air, from water, or from deposits ...

Eva Koppelhus.

Classification

''Unescoceratops'' has been classified as a

and the sister taxa to ''

Gryphoceratops ''Gryphoceratops'' is an extinct genus of leptoceratopsid ceratopsian dinosaur known from the Late Cretaceous of Alberta, southern Canada. Discovery ''Gryphoceratops'' is known only from the holotype ROM 56635, a partial right dentary. Th ...

'', which was described in the same paper. These two are the most derived leptoceratopsids known, despite ''Gryphoceratops'' also being the oldest leptoceratopsid. The cladogram below represents the cladistic relationship of ''Unescoceratops'' compared to other

leptoceratopsids Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but ...

as reconstructed from the phylogenetic analysis.

References

Fossils of Canada Leptoceratopsids Late Cretaceous dinosaurs of North America Fossil taxa described in 2012 Taxa named by Philip J. Currie Dinosaur Park fauna Paleontology in Alberta Campanian genus first appearances Campanian genus extinctions Ornithischian genera {{ceratopsian-stub

Discovery

Description

Etymology

Classification

See also

References