Thalassemys
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''Thalassemys'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including '' Eurysternum'', ''Plesiochelys'' and '' Thalassemys'' to the exclusion of '' Pe ...
n
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked t ...
from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
of western and central
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. While the genus was originally named by Rütimeyer in 1859 for a large
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
and other associated fragments from the late
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
of the
Reuchenette Formation The Reuchenette Formation is a Jurassic geologic formation in Switzerland. It is Kimmeridgian in age and predominantly consists of well stratified limestone, with lithology variable both laterally and stratigraphically including wackestones, pac ...
of
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
, although the taxon was not validly named until 1873 when Rütimeyer designated the type species ''T. hugii''. Rütimeyer also named ''T. gresslyi'' from the Reunchenette Formation in the same paper as ''T. hugii'', but it cannot be differentiated from the type material of ''T. hugii'' and is therefore a junior synonym. A large assemblage of shell and postcranial material from the Reunchenette was named as a species of '' Eurysternum'', ''E. ignoratum'', by Bräm in
1965 Events January–February * January 14 – The Prime Minister of Northern Ireland and the Taoiseach of the Republic of Ireland meet for the first time in 43 years. * January 20 ** Lyndon B. Johnson is sworn in for a full term ...
. While originally distinguished based on the presence of fontanelles on the plastron, the feature was later identified on ''T. hugii'' and ''E. ignoratum'' was designated a junior synonym. Additional material from the
Kimmeridge Clay The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for Nor ...
of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and ...
has also been referred to ''T. hugii''. The second valid species of ''Thalassemys'' is ''T. marina'', named in
1903 Events January * January 1 – Edward VII is proclaimed Emperor of India. * January 19 – The first west–east transatlantic radio broadcast is made from the United States to England (the first east–west broadcast having bee ...
by Fraas for a partial carapace and half a plastron from the late Kimmeridgian deposits of
Schnaitheim Heidenheim an der Brenz, or just Heidenheim (; Swabian: ''Hoidna'' or ''Hoirna''), is a town in Baden-Württemberg in southern Germany. It is located near the border with Bavaria, approximately 17 km south of Aalen and 33 km north of ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwee ...
. It was suggested to be a species of the related taxon '' Eurysternum'' by Maisch in
2001 The September 11 attacks against the United States by Al-Qaeda, which killed 2,977 people and instigated the global war on terror, were a defining event of 2001. The United States led a multi-national coalition in an invasion of Afghanist ...
, who referred additional material from Schnaitheim to the species. In
2003 File:2003 Events Collage.png, From top left, clockwise: The crew of STS-107 perished when the Space Shuttle Columbia disintegrated during reentry into Earth's atmosphere; SARS became an epidemic in China, and was a precursor to SARS-CoV-2; A ...
, it was suggested that ''T. marina'' was a synonym of the related turtle '' Palaeomedusa'', which was based on an unknown but restored region of the original carapace. ''Thalassemys heusseri'' was validly named in
1924 Events January * January 12 – Gopinath Saha shoots Ernest Day, whom he has mistaken for Sir Charles Tegart, the police commissioner of Calcutta, and is arrested soon after. * January 20– 30 – Kuomintang in China hold ...
by Oertel for a partial carapace from the Kimmeridgian or
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by ...
of Holzen, Germany, a different spelling from the invalid name "T. heuseri" he used previously in
1815 Events January * January 2 – Lord Byron marries Anna Isabella Milbanke in Seaham, county of Durham, England. * January 3 – Austria, Britain, and Bourbon-restored France form a secret defensive alliance treaty against Pru ...
. As the type specimen was lost, the species cannot be distinguished from others of the genus, and is therefore a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
''. In addition to ''E. ignoratum'', Bräm also named the species ''Thalassemys moseri'' in 1965 for a partial carapace from the Reunchenette Formation. An additional shell and also a skull were referred to the taxon in
1980 Events January * January 4 – U.S. President Jimmy Carter proclaims a grain embargo against the USSR with the support of the European Commission. * January 6 – Global Positioning System time epoch begins at 00:00 UTC. * January 9 – In ...
by Rieppel, who suggested that the species should be included in '' Plesiochelys'' rather than ''Thalassemys''. This skull was given the name '' Jurassichelon'' in
2015 File:2015 Events Collage new.png, From top left, clockwise: Civil service in remembrance of November 2015 Paris attacks; Germanwings Flight 9525 was purposely crashed into the French Alps; the rubble of residences in Kathmandu following the April ...
, leaving the species ''T. moseri'', based on other material, in an uncertain position. Further assessment of the material by Anquetin and colleagues in
2017 File:2017 Events Collage V2.png, From top left, clockwise: The War Against ISIS at the Battle of Mosul (2016-2017); aftermath of the Manchester Arena bombing; The Solar eclipse of August 21, 2017 ("Great American Eclipse"); North Korea tests a s ...
concluded that ''T. moseri'' was distinct from the species ''J. oleronensis'' named for the skull, but was representative of the same genus. ''Thalassemys bruntrutana'' was described in 2015 by Püntener and colleagues for an almost complete carapace and plastron from a lower member of the Reunchenette Formation than other species, with referred material also from the British
Kimmeridge Clay The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for Nor ...
of the
Isle of Purbeck The Isle of Purbeck is a peninsula in Dorset, England. It is bordered by water on three sides: the English Channel to the south and east, where steep cliffs fall to the sea; and by the marshy lands of the River Frome and Poole Harbour to the no ...
. While the genus ''Thalassemys'' was placed in the family Thalassemydidae, it represents the only definitive member of the family due to the uncertain relationships of other members of
Thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including '' Eurysternum'', ''Plesiochelys'' and '' Thalassemys'' to the exclusion of '' Pe ...
, primarily due to the lack of cranial material in ''Thalassemys''.


References

* {{Taxonbar, from=Q7709531 Thalassochelydia Prehistoric turtle genera Kimmeridgian genera Late Jurassic turtles Late Jurassic reptiles of Europe Jurassic England Fossils of England Jurassic Germany Fossils of Germany Jurassic Switzerland Fossils of Switzerland Fossil taxa described in 1873