Tessellated darter
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The tessellated darter (''Etheostoma olmstedi'') is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a darter from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Etheostomatinae Etheosomatidae is a species rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family i ...
, part of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more than ...
, which also contains the
perch Perch is a common name for fish of the genus ''Perca'', freshwater gamefish belonging to the family Percidae. The perch, of which three species occur in different geographical areas, lend their name to a large order of vertebrates: the Percif ...
es, ruffes and pikeperches. It is native to Canada and the United States. It was formerly considered a subspecies of the johnny darter, which it greatly resembles in general appearance.


Description

Both the tessellated darter and johnny darter are rather pale with small X or W shaped markings on the back and upper sides. Both have only a single anal fin spine whereas all other darters in their range have two. Tessellated darters have somewhat sharper snouts than johnny darters, more dorsal soft rays, and more pectoral fin rays. The suborbital sensory canal is complete in most tessellated darters, usually broken into two sections in johnny darters. Their nape is usually naked, and their cheeks are partly to completely scaled. The breast is usually naked and the belly partly scaled. The body of the tessellated darter is elongate, but a little compressed. In breeding males, the spines have fleshy knobs although they are not as well developed as in the fantail darter. They are a pale, sandy color shading to whitish along the body ventrally. Some scales of the upper sides have dark margins forming wavy bars of varying lengths and different angles. Their sides have X and W shaped marks, of which 9 to 11 are prominent along the midside. Dorsal fins have a dark pigment spot on the first interradial membrane, and the rest of the fin is clear or slightly dusky. the snout has a dark line from each eye to the nostrils. Juveniles and breeding adults develop 12 or 13 rather even vertical bands along their sides while losing the wavy lines and X shaped markings as the scales of the upper sides become quite dark with pale tips on the pectoral and pelvic fins. The unpigmented areas of the second dorsal, anal, and caudal fins stand out so that the fin has the appearance of having white bands on a dark background, which is a reversal of the appearance in non-breeding adults. In this color phase, they are sometimes mistaken for a species of ''
Percina ''Percina'' is a genus of small freshwater ray-finned fish, classified within the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches from North America. Like the similar fishes in the ...
''.


Ecology

Tessellated darters eat crustaceans and small insects when they are small, gradually shifting to larger insects as the fish get bigger. Male tessellated darters guard nests of fertilized eggs until the fry (young) are free-swimming, and have been observed to engage in alloparental (adoptive) care of previous nest inhabitants' eggs. Alloparental care is associated with increased male reproductive success in this species. Males frequently engage in filial cannibalism (consumption of their own offspring).


Taxonomy and etymology

The tessellated darter was first formally described in 1842 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
David Humphreys Storer with the type locality given as the
Connecticut River The Connecticut River is the longest river in the New England region of the United States, flowing roughly southward for through four states. It rises 300 yards (270 m) south of the U.S. border with Quebec, Canada, and discharges at Long Island ...
at
Hartford, Connecticut Hartford is the capital city of the U.S. state of Connecticut. It was the seat of Hartford County until Connecticut disbanded county government in 1960. It is the core city in the Greater Hartford metropolitan area. Census estimates since the ...
. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honors the collector of the type, Charles H. Olmsted who was president of the Hartford Natural History Society. It is closely related to the Johnny darter ''E. nigrum'' and has been considered
synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
with it.


References

*"The Inland Fish of New York State." C. Lavett Smith. olmstedi Taxa named by David Humphreys Storer Fish described in 1842 {{Percidae-stub