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A reef is a ridge or
shoal In oceanography, geomorphology, and geoscience, a shoal is a natural submerged ridge, bank, or bar that consists of, or is covered by, sand or other unconsolidated material and rises from the bed of a body of water to near the surface. It ...
of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock outcrops, etc.—but there are also reefs such as the coral reefs of tropical waters formed by
biotic Biotics describe living or once living components of a community; for example organisms, such as animals and plants. Biotic may refer to: *Life, the condition of living organisms *Biology, the study of life * Biotic material, which is derived from ...
processes dominated by corals and coralline algae, and artificial reefs such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, to attract a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this. Earth's largest coral reef system is the Great Barrier Reef in Australia, at a length of over .


Biotic

There is a variety of biotic reef types, including oyster reefs and sponge reefs, but the most massive and widely distributed are tropical coral reefs. Although corals are major contributors to the framework and bulk material comprising a coral reef; the organisms most responsible for reef growth against the constant assault from ocean waves are calcareous algae, especially, although not entirely, coralline algae. The preferred substrate for oyster larvae is the shells of oysters so they tend to settle on adult oysters and thereby develop layers building upwards, eventually forming a fairly massive hard stony calcium carbonate structure on which other reef organisms like
sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through ...
and seaweeds can grow, and provide a habitat for mobile benthic organisms. These biotic reef types take on additional names depending upon how the reef lies in relation to the land, if any. Reef types include fringing reefs, barrier reefs, and atolls. A fringing reef is a reef that is attached to an island. A barrier reef forms a calcareous barrier around an island, resulting in a lagoon between the shore and the reef. An atoll is a ring reef with no land present. The reef front (ocean side) is a high energy locale, whereas the internal lagoon will be at a lower energy with fine grained sediments.


Mounds

One definition distinguishes reefs from mounds as follows: both are considered to be varieties of organosedimentary buildups – sedimentary features, built by the interaction of organisms and their environment, that have synoptic relief and whose biotic composition differs from that found on and beneath the surrounding
sea floor The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean. All floors of the ocean are known as 'seabeds'. The structure of the seabed of the global ocean is governed by plate tectonics. Most of ...
. Reefs are held up by a macroscopic skeletal framework. Coral reefs are an example of this kind. Corals and calcareous algae grow on top of one another and form a three-dimensional framework that is modified in various ways by other organisms and inorganic processes. By contrast, mounds lack a macroscopic skeletal framework. Mounds are built by microorganisms or by organisms that don't grow a skeletal framework. A microbial mound might be built exclusively or primarily by
cyanobacteria Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, blu ...
. Examples of
biostrome A reef knoll is a land-based landform that comprises an immense pile of calcareous material that accumulated on a previously existing ancient sea floor. At the time of its accumulation it may have had enough structure from organisms such as sponges ...
s formed by cyanobacteria occur in the
Great Salt Lake The Great Salt Lake is the largest saltwater lake in the Western Hemisphere and the eighth-largest terminal lake in the world. It lies in the northern part of the U.S. state of Utah and has a substantial impact upon the local climate, particula ...
in Utah, United States, and in
Shark Bay Shark Bay (Malgana: ''Gathaagudu'', "two waters") is a World Heritage Site in the Gascoyne region of Western Australia. The http://www.environment.gov.au/heritage/places/world/shark-bay area is located approximately north of Perth, on the ...
on the coast of Western Australia. Cyanobacteria do not have skeletons, and individuals are microscopic. Cyanobacteria can encourage the precipitation or accumulation of calcium carbonate to produce distinct sediment bodies in composition that have relief on the seafloor. Cyanobacterial mounds were most abundant before the evolution of shelly macroscopic organisms, but they still exist today; stromatolites are microbial mounds with a laminated internal structure. Bryozoans and crinoids, common contributors to marine sediments during the
Mississippian Mississippian may refer to: * Mississippian (geology), a subperiod of the Carboniferous period in the geologic timescale, roughly 360 to 325 million years ago *Mississippian culture, a culture of Native American mound-builders from 900 to 1500 AD ...
, for instance, produced a very different kind of mound. Bryozoans are small and the skeletons of crinoids disintegrate. However, bryozoan and crinoid meadows can persist over time and produce compositionally distinct bodies of sediment with depositional relief. The
Proterozoic The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...
Belt Supergroup The Belt Supergroup is an assemblage of primarily fine-grained sedimentary rocks and mafic intrusive rocks of late Precambrian (Mesoproterozoic) age. It is more than thick, covers an area of some 200,000 km2 (77,220 sq. mi), and is considered ...
contains evidence of possible microbial mat and dome structures similar to stromatolite and chicken reef complexes.


Geologic

Rocky reefs are underwater outcrops of rock projecting above the adjacent unconsolidated surface with varying relief. They can be found in depth ranges from intertidal to deep water, and provide a substrate for a large range of sessile benthic organisms, and shelter for a large range of mobile organisms.


Structures

Ancient reefs buried within stratigraphic sections are of considerable interest to geologists because they provide paleo-environmental information about the location in
Earth's history The history of Earth concerns the development of planet Earth from its formation to the present day. Nearly all branches of natural science have contributed to understanding of the main events of Earth's past, characterized by constant geologic ...
. In addition, reef structures within a sequence of sedimentary rocks provide a discontinuity which may serve as a trap or conduit for
fossil fuel A fossil fuel is a hydrocarbon-containing material formed naturally in the Earth's crust from the remains of dead plants and animals that is extracted and burned as a fuel. The main fossil fuels are coal, oil, and natural gas. Fossil fuels m ...
s or mineralizing fluids to form petroleum or ore deposits. Corals, including some major extinct groups Rugosa and Tabulata, have been important reef builders through much of the
Phanerozoic The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
since the Ordovician Period. However, other organism groups, such as calcifying algae, especially members of the red algae (
Rhodophyta Red algae, or Rhodophyta (, ; ), are one of the oldest groups of eukaryotic algae. The Rhodophyta also comprises one of the largest phyla of algae, containing over 7,000 currently recognized species with taxonomic revisions ongoing. The majority ...
), and molluscs (especially the rudist bivalves during the Cretaceous Period) have created massive structures at various times. During the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
Period, the conical or tubular skeletons of Archaeocyatha, an extinct group of uncertain affinities (possibly sponges), built reefs. Other groups, such as the Bryozoa, have been important interstitial organisms, living between the framework builders. The corals which build reefs today, the Scleractinia, arose after the Permian–Triassic extinction event that wiped out the earlier rugose corals (as well as many other groups), and became increasingly important reef builders throughout the Mesozoic Era. They may have arisen from a rugose coral ancestor. Rugose corals built their skeletons of
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
and have a different symmetry from that of the scleractinian corals, whose skeletons are aragonite. However, there are some unusual examples of well-preserved aragonitic rugose corals in the Late Permian. In addition, calcite has been reported in the initial post-larval calcification in a few scleractinian corals. Nevertheless, scleractinian corals (which arose in the middle Triassic) may have arisen from a non-calcifying ancestor independent of the rugosan corals (which disappeared in the late Permian).


Artificial

An artificial reef is a human-created underwater structure, typically built to promote
marine life Marine life, sea life, or ocean life is the plants, animals and other organisms that live in the salt water of seas or oceans, or the brackish water of coastal estuaries. At a fundamental level, marine life affects the nature of the planet. M ...
in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of trawling nets, or improve
surfing Surfing is a surface water sport in which an individual, a surfer (or two in tandem surfing), uses a board to ride on the forward section, or face, of a moving wave of water, which usually carries the surfer towards the shore. Waves suitabl ...
. Many reefs are built using objects that were built for other purposes, for example by sinking oil rigs (through the Rigs-to-Reefs program), scuttling ships, or by deploying rubble or construction debris. Other artificial reefs are purpose built (e.g. the
reef balls Reef Ball Foundation, Inc. is a 501(c)(3) non-profit organization that functions as an international environmental non-governmental organization. The foundation uses reef ball artificial reef technology, combined with coral propagation, transpla ...
) from PVC or concrete. Shipwrecks become artificial reefs on the seafloor. Regardless of construction method, artificial reefs generally provide stable hard surfaces where
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
and invertebrates such as
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in eros ...
s, corals, and oysters attach; the accumulation of attached marine life in turn provides intricate structure and food for assemblages of fish.


See also

* * * *


References


Sources

* Shears N.T. (2007) Biogeography, community structure and biological habitat types of subtidal reefs on the South Island West Coast, New Zealand. ''Science for Conservation 281''. p 53. Department of Conservation, New Zealand


External links


''Reef Rescue'' - Smithsonian Ocean Portal

Coral Reefs of the Tropics
: facts, photos and movies from The Nature Conservancy
NOAA Photo Library

Reef Environmental Education Foundation

NOS Data Explorer
- A portal to obtain NOAA National Ocean Service data


Atolls – Distribution, Development and Architecture
{{Authority control * Stratigraphy Coastal geography Coastal and oceanic landforms