Syrian wild ass
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The Syrian wild ass (''Equus hemionus hemippus''), less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of
onager The onager (; ''Equus hemionus'' ), A new species called the kiang (''E. kiang''), a Tibetan relative, was previously considered to be a subspecies of the onager as ''E. hemionus kiang'', but recent molecular studies indicate it to be a distinct ...
native to the
Arabian peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
and surrounding areas. It ranged across present-day
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
,
Palestine __NOTOC__ Palestine may refer to: * State of Palestine, a state in Western Asia * Palestine (region), a geographic region in Western Asia * Palestinian territories, territories occupied by Israel since 1967, namely the West Bank (including East ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
, and
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
.


Description

The Syrian wild ass, one metre high at its shoulder, was the smallest
equine Equinae is a subfamily of the family Equidae, which have lived worldwide (except Indonesia and Australia) from the Hemingfordian stage of the Early Miocene (16 million years ago) onwards. They are thought to be a monophyletic grouping.B. J. MacFa ...
, and it could not be domesticated. Its coloring changed with the seasons — a tawny olive coat for the summer months, and pale sandy yellow for the winter. It was known, like other onagers, to be untameable, and was compared to a thoroughbred horse for its beauty and strength.


Distribution and habitat

The Syrian wild ass lived in deserts, semi-deserts, arid grasslands, and mountain steppes. Native to
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
, they were found in
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Palestine __NOTOC__ Palestine may refer to: * State of Palestine, a state in Western Asia * Palestine (region), a geographic region in Western Asia * Palestinian territories, territories occupied by Israel since 1967, namely the West Bank (including East ...
, Jordan, Turkey, Syria, Saudi Arabia, and
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
.


Ecology and behavior


Diet

The Syrian wild ass was a
grazer Grazer may refer to: *grazer, an animal that grazes *Grazer, a native or inhabitant of Graz *GRAZER, the shoegaze band GRAZER Astronomy * Earth Grazer, Earth-grazing fireball that enters the Earth's atmosphere and leaves again * Mercury grazer, ...
and a browser. It fed on grass, herbs, leaves, shrubs, and tree branches.


Predation

Syrian wild asses were preyed upon by
Asiatic lion The Asiatic lion is a population of ''Panthera leo leo'' that today survives in the wild only in India. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of Gujarat ...
s,
Arabian leopard The Arabian leopard (''Panthera pardus nimr'') is a leopard subspecies native to the Arabian Peninsula. It has been listed as Critically Endangered on the IUCN Red List since 1996 as fewer than 200 wild individuals were estimated to be alive i ...
s, striped hyenas, Syrian brown bears, Arabian wolves, and
Caspian tiger The Caspian tiger was a ''Panthera tigris tigris'' population native to eastern Turkey, northern Iran, Mesopotamia, the Caucasus around the Caspian Sea, Central Asia to northern Afghanistan, and the Xinjiang region in western China. Until the Midd ...
s.
Asiatic cheetah The Asiatic cheetah (''Acinonyx jubatus venaticus'') is a critically endangered cheetah subspecies currently only surviving in Iran. It once occurred from the Arabian Peninsula and the Near East to the Caspian region, Transcaucasus, Kyzylkum D ...
s and
golden jackal The golden jackal (''Canis aureus''), also called common jackal, is a wolf-like canid that is native to Southeast Europe, Southwest Asia, South Asia, and regions of Southeast Asia. The golden jackal's coat varies in color from a pale creamy y ...
s may have also preyed on foals.


Relationship with humans

The bones of a Syrian wild ass have been identified at an 11,000 year-old archaeological site at
Göbekli Tepe Göbekli Tepe (, "Potbelly Hill"; known as ''Girê Mirazan'' or ''Xirabreşkê'' in Kurdish languages, Kurdish) is a Neolithic archaeological site in the Southeastern Anatolia Region of Turkey. Dated to the Pre-Pottery Neolithic, between 9500 ...
, Turkey. Cuneiform from the third millennium  report the hunting of an 'equid of the desert' (''anše-edin-na''), valued for its meat and hide, which may have been ''E.h. hemippus''. Although Syrian wild asses were not themselves domesticated, a significant breeding center at
Tell Brak Tell Brak (Nagar, Nawar) was an ancient city in Syria; its remains constitute a tell located in the Upper Khabur region, near the modern village of Tell Brak, 50 kilometers north-east of Al-Hasaka city, Al-Hasakah Governorate. The city' ...
produced a hybrid of the wild ass and the donkey, called the '' kunga'', that was a draft animal of high economic and symbolic value to the elite of Syria and Mesopotamia. They appear in cuneiform inscriptions and their bones are found in burials from the third millennium . The size of these hybrids, larger than modern examples of both parent species, has led to speculation that the Syrian wild asses used historically in breeding the kunga were of larger size than the individuals observed in the remant populations of the 18th and 19th centuries.
Assyria Assyria (Neo-Assyrian cuneiform: , romanized: ''māt Aššur''; syc, ܐܬܘܪ, ʾāthor) was a major ancient Mesopotamian civilization which existed as a city-state at times controlling regional territories in the indigenous lands of the A ...
n art from the 7th century  found at
Nineveh Nineveh (; akk, ; Biblical Hebrew: '; ar, نَيْنَوَىٰ '; syr, ܢܝܼܢܘܹܐ, Nīnwē) was an ancient Assyrian city of Upper Mesopotamia, located in the modern-day city of Mosul in northern Iraq. It is located on the eastern ban ...
includes a scene of hunters capturing Syrian wild asses with
lasso A lasso ( or ), also called lariat, riata, or reata (all from Castilian, la reata 're-tied rope'), is a loop of rope designed as a restraint to be thrown around a target and tightened when pulled. It is a well-known tool of the Spanish an ...
s.
Xenophon of Athens Xenophon of Athens (; grc, Ξενοφῶν ; – probably 355 or 354 BC) was a Greek military leader, philosopher, and historian, born in Athens. At the age of 30, Xenophon was elected commander of one of the biggest Greek mercenary armies of ...
mentions Syrian wild asses in his ''
Anabasis Anabasis (from Greek ''ana'' = "upward", ''bainein'' = "to step or march") is an expedition from a coastline into the interior of a country. Anabase and Anabasis may also refer to: History * ''Anabasis Alexandri'' (''Anabasis of Alexander''), a ...
'' of ~370 . He reports that they were the most common of animals encountered in Syria; in addition to
ostriches Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There are ...
,
bustards Bustards, including floricans and korhaans, are large, terrestrial birds living mainly in dry grassland areas and on the steppes of the Old World. They range in length from . They make up the family Otididae (, formerly known as Otidae). Bust ...
, and gazelles. Xenophon states that horsemen would occasionally chase the asses, with the asses easily able to outrun the horses. He said that asses would only run a short distance ahead of the horses before stopping, waiting for the horses to get closer, and then running ahead yet again. He described the asses as impossible to catch without careful planning. Xenophon also related that the meat of the asses tasted like a more tender version of venison. It is believed this may be the "wild ass" that
Ishmael Ishmael ''Ismaḗl''; Classical/Qur'anic Arabic: إِسْمَٰعِيْل; Modern Standard Arabic: إِسْمَاعِيْل ''ʾIsmāʿīl''; la, Ismael was the first son of Abraham, the common patriarch of the Abrahamic religions; and is cons ...
was prophesied to be in
Genesis Genesis may refer to: Bible * Book of Genesis, the first book of the biblical scriptures of both Judaism and Christianity, describing the creation of the Earth and of mankind * Genesis creation narrative, the first several chapters of the Book of ...
in the
Old Testament The Old Testament (often abbreviated OT) is the first division of the Christian biblical canon, which is based primarily upon the 24 books of the Hebrew Bible or Tanakh, a collection of ancient religious Hebrew writings by the Israelites. The ...
. References also appear in the Old Testament books of
Job Work or labor (or labour in British English) is intentional activity people perform to support the needs and wants of themselves, others, or a wider community. In the context of economics, work can be viewed as the human activity that contr ...
,
Psalms The Book of Psalms ( or ; he, תְּהִלִּים, , lit. "praises"), also known as the Psalms, or the Psalter, is the first book of the ("Writings"), the third section of the Tanakh, and a book of the Old Testament. The title is derived ...
,
Jeremiah Jeremiah, Modern:   , Tiberian: ; el, Ἰερεμίας, Ieremíās; meaning " Yah shall raise" (c. 650 – c. 570 BC), also called Jeremias or the "weeping prophet", was one of the major prophets of the Hebrew Bible. According to Jewish ...
, and the Deuterocanonical book of
Sirach The Book of Sirach () or Ecclesiasticus (; abbreviated Ecclus.) is a Jewish work, originally in Hebrew, of ethical teachings, from approximately 200 to 175 BC, written by the Judahite scribe Ben Sira of Jerusalem, on the inspiration of his fa ...
. The
Qur’an The Quran (, ; Standard Arabic: , Quranic Arabic: , , 'the recitation'), also romanized Qur'an or Koran, is the central religious text of Islam, believed by Muslims to be a revelation from God. It is organized in 114 chapters (pl.: , sing.: ...
, the main book of
Islam Islam (; ar, ۘالِإسلَام, , ) is an Abrahamic religions, Abrahamic Monotheism#Islam, monotheistic religion centred primarily around the Quran, a religious text considered by Muslims to be the direct word of God in Islam, God (or ...
, in Surat al-Muddaththir, refers to a scene of ''humur'' ( ar, حُـمـر, 'asses' or 'donkeys' in plural form, حمار singular) fleeing from a ''qaswarah'' ( ar, قَـسـورة, '
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphi ...
'). This was to criticize people who were averse to
Muhammad Muhammad ( ar, مُحَمَّد;  570 – 8 June 632 Common Era, CE) was an Arab religious, social, and political leader and the founder of Islam. According to Muhammad in Islam, Islamic doctrine, he was a prophet Divine inspiration, di ...
's teachings, such as supporting the welfare of the less wealthy.


Later hybrids

In addition to the Bronze Age kunga, a couple of modern hybrids were produced by the
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for science, scientific study. In 1831 o ...
in the late 19th century. In 1878, a Syrian wild ass was crossed with an
Indian wild ass The Indian wild ass (''Equus hemionus khur''), also called the Indian onager or, in the local Gujarati language, Ghudkhur and Khur, is a subspecies of the onager native to South Asia. It is currently listed as Near Threatened by IUCN. The prev ...
(a different subspecies), and in 1883 an inter-species cross between a Syrian wild ass male and an Abyssinian wild ass female produced a foal that was colored like the sire, and described as "a fine animal" but "vicious and untamed".


Extinction

European travelers in the Middle East during the fifteenth and sixteenth centuries reported seeing large herds. However, its numbers began to drop precipitously during the eighteenth and nineteenth centuries due to overhunting, and its existence was further imperiled by the regional upheaval of
World War I World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
. The last known wild specimen was fatally shot in 1927 at al Ghams near the
Azraq oasis Azraq ( ar, الأزرق meaning "blue") is a small town in Zarqa Governorate in central-eastern Jordan, east of Amman. The population of Azraq was 9,021 in 2004. The Muwaffaq Salti Air Base is located in Azraq. History Prehistory Archaeo ...
in Jordan, and the last captive specimen died the same year at the
Tiergarten Schönbrunn Tiergarten (German for "zoo") may refer to: * Tiergarten (park), an urban public park in Berlin * Kleiner Tiergarten, a smaller public park in Berlin * Tiergarten, Berlin, a locality within the borough of Mitte which includes the park * Berlin-Tie ...
, in Vienna.


Replacement

After the extinction of the Syrian wild ass, the
Persian onager The Persian onager (''Equus hemionus onager''), also called the Persian wild ass or Persian zebra, is a subspecies of onager (Asiatic wild ass) native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wil ...
from
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
was chosen as an appropriate subspecies to repopulate the Middle East as a replacement for the extinct ''E. h. hemippus'' onagers. The Persian onager was then introduced to the protected areas of Saudi Arabia and Jordan. It also was reintroduced, along with the
Turkmenian kulan The Turkmenian kulan (''Equus hemionus kulan''), also called Transcaspian wild ass, Turkmenistani onager or simply the ', is a subspecies of onager (Asiatic wild ass) native to Central Asia. It was declared Endangered in 2016. The species's popu ...
, to Israel, where they both reproduce wild ass hybrids in the
Negev Mountains Negev Mountains is a mountainous area in the north-western part of the Negev desert, in Israel. Mount Ramon is the summit of Negev Mountains and the highest point in southern Israel, reaching . Most of the area belongs to Negev Mountains Nature ...
and the
Yotvata Hai-Bar Nature Reserve The Yotvata Hai-Bar Nature Reserve is a breeding and reacclimation center administered by the Israel Nature Reserves & National Parks Authority, situated in the Southern Arava near Yotvata. The Yotvata Hai-Bar is the desert counterpart of t ...
.


Related subspecies

*
Mongolian wild ass The Mongolian wild ass (''Equus hemionus hemionus''), also known as Mongolian khulan, is the nominate subspecies of the onager. It is found in southern Mongolia and northern China. It was previously found in eastern Kazakhstan and southern Siberi ...
(''khulan''), ''Equus hemionus hemionus'' *
Turkmenian kulan The Turkmenian kulan (''Equus hemionus kulan''), also called Transcaspian wild ass, Turkmenistani onager or simply the ', is a subspecies of onager (Asiatic wild ass) native to Central Asia. It was declared Endangered in 2016. The species's popu ...
(''kulan''), ''Equus hemionus kulan'' *
Indian wild ass The Indian wild ass (''Equus hemionus khur''), also called the Indian onager or, in the local Gujarati language, Ghudkhur and Khur, is a subspecies of the onager native to South Asia. It is currently listed as Near Threatened by IUCN. The prev ...
(''khur''), ''Equus hemionus khur'' *
Persian onager The Persian onager (''Equus hemionus onager''), also called the Persian wild ass or Persian zebra, is a subspecies of onager (Asiatic wild ass) native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wil ...
(''gur''), ''Equus hemionus onager''


References

{{Taxonbar, from=Q946292 1927 in the environment Onager Extinct mammals of Asia Mammal extinctions since 1500 Mammals described in 1855 Species endangered by warfare Species made extinct by human activities