Subalpine fir
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''Abies lasiocarpa'', the subalpine fir or Rocky Mountain fir, is a western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
n fir tree.


Description

''Abies lasiocarpa'' is a medium-sized
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
conifer Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ext ...
with a very narrow conic crown, growing to tall, exceptionally , with a
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
up to thick, exceptionally . The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
on young trees is smooth, gray, and with
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...
blisters, becoming rough and fissured or scaly on old trees. The lowest branches can be observed growing above ground level. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are flat and needle-like, long, glaucous green above with a broad stripe of
stomata In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is b ...
, and two blue-white stomatal bands below; the fresh leaf scars are reddish (tan on the inland variety). They are arranged spirally on the shoot, but with the leaf bases twisted to be arranged to the sides of and above the shoot, with few or none below the shoot. The cones are erect, long, dark purple with fine yellow-brown pubescence, ripening brown and disintegrating to release the winged
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s in early fall.


Taxonomy

There are two or three
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
in subalpine fir, treated very differently by different authors: *The Coast Range subalpine fir (''Abies lasiocarpa'') in the narrow sense, is the typical form of the species, occurring in the
Pacific Coast Ranges The Pacific Coast Ranges (officially gazetted as the Pacific Mountain System in the United States) are the series of mountain ranges that stretch along the West Coast of North America from Alaska south to Northern and Central Mexico. Although the ...
, the
Olympic Mountains The Olympic Mountains are a mountain range on the Olympic Peninsula of the Pacific Northwest of the United States. The mountains, part of the Pacific Coast Ranges, are not especially high – Mount Olympus is the highest at ; however, the easte ...
and the Cascade Range from Southeast Alaska (Panhandle mountains) south to California. *The Rocky Mountains subalpine fir is very closely related and of disputed status, being variously treated as a distinct species ''Abies bifolia'', as a
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
of Coast Range subalpine fir ''Abies lasiocarpa'' var. ''bifolia'', or not distinguished from typical ''A. lasiocarpa'' at all. It occurs in the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
from Southeast Alaska (eastern
Alaska Range The Alaska Range is a relatively narrow, 600-mile-long (950 km) mountain range in the southcentral region of the U.S. state of Alaska, from Lake Clark at its southwest endSources differ as to the exact delineation of the Alaska Range. ThBo ...
) south to Colorado. It differs primarily in the chemical composition of its resin, microscopic features, and in the fresh leaf scars being yellow-brown, not reddish. The ''Flora of North America'' treats it as a distinct species, while the
United States Department of Agriculture The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of comme ...
(USDA) includes it within ''A. lasiocarpa'' without distinction. *The corkbark fir ''Abies lasiocarpa'' var. ''arizonica'' occurs in Arizona and New Mexico. It differs in thicker, corky bark and more strongly glaucous foliage. In resin composition it is closer to ''A. bifolia'' than to typical ''A. lasiocarpa'', though the combination "Abies bifolia var. arizonica" has not been formally published. The ''Flora of North America'' includes it within ''A. bifolia'' without distinction; the USDA treats it as a distinct variety of ''A. lasiocarpa''.


Distribution

The species is native to the mountains of
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
,
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
and western
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest T ...
in
Western Canada Western Canada, also referred to as the Western provinces, Canadian West or the Western provinces of Canada, and commonly known within Canada as the West, is a Canadian region that includes the four western provinces just north of the Canadaâ ...
; and to
Southeast Alaska Southeast Alaska, colloquially referred to as the Alaska(n) Panhandle, is the southeastern portion of the U.S. state of Alaska, bordered to the east and north by the northern half of the Canadian province of British Columbia (and a small part ...
,
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
,
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
,
Idaho Idaho ( ) is a U.S. state, state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Monta ...
,
Western Montana Western Montana is the western region of the U.S. state of Montana. The most restrictive definition limits western Montana only to the parts of the state west of the Continental Divide. Other common definitions add in the mountainous areas east ...
,
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to t ...
,
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
,
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
,
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, AlÄ­ á¹£onak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
, northeastern
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
, and the Trinity Alps of the
Klamath Mountains The Klamath Mountains are a rugged and lightly populated mountain range in northwestern California and southwestern Oregon in the western United States. As a mountain system within both the greater Pacific Coast Ranges and the California Coast ...
in northwestern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
in the
Western United States The Western United States (also called the American West, the Far West, and the West) is the region comprising the westernmost states of the United States. As American settlement in the U.S. expanded westward, the meaning of the term ''the We ...
. It occurs at high altitudes, commonly found at and immediately below the
tree line The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snow ...
. It can be found at elevations of in the north of the range (rarely down to sea level in the far north), to in the south of the range. West of the
Cascade Cascade, Cascades or Cascading may refer to: Science and technology Science *Cascade waterfalls, or series of waterfalls * Cascade, the CRISPR-associated complex for antiviral defense (a protein complex) * Cascade (grape), a type of fruit * Bioc ...
ridge, it can be found at elevations of , while further east (particularly in Western Montana) it can be found from .


Ecology

Annual precipitation ranges from in coastal mountain sites to only inland. Snow gathered on the branches helps protect them from wind and heat. Firs in general act as a
snow fence A snow fence, similar to a sand fence, is a barrier that forces windblown, drifting snow to accumulate in a desired place. They are primarily employed to minimize the amount of snowdrift on roadways and railways. Farmers and ranchers use snow ...
, leading to the creation of meadows through extra moisture accumulation. The tree is highly
shade tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial ...
, but very vulnerable to fire, short-lived, and slow-growing. Despite having weaker wood than some of its timberline associates, it can survive by its ability to adapt (growing in a ''
krummholz ''Krummholz'' (german: krumm, "crooked, bent, twisted" and ''Holz'', "wood") — also called ''knieholz'' ("knee timber") — is a type of stunted, deformed vegetation encountered in the subarctic and subalpine tree line landscapes, shaped b ...
'' form) and reproduce via
layering Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...
in clusters at high elevations. At timberline, a single tree can leave behind a ring of trees (an 'atoll') via layering. The species has benefited from
wildfire suppression Wildfire suppression is a range of firefighting tactics used to suppress wildfires. Firefighting efforts in wild land areas require different techniques, equipment, and training from the more familiar structure fire fighting found in populated ...
in more recent years. Various animals, including mountain goats, take shelter in subalpine fir clusters and ''krummholz''. The bark is browsed by game animals and its leaves are eaten by grouse. Songbirds, Richardson's grouse, Cascade pine squirrels, and other mammals consume the seeds. It is host to
pathogenic fungi Pathogenic fungi are fungi that cause disease in humans or other organisms. Approximately 300 fungi are known to be pathogenic to humans. Markedly more fungi are known to be pathogenic to plant life than those of the animal kingdom. The study of ...
such as the species '' Delphinella balsameae''.


Uses

Native Americans used the leaves as deodorant and burned them as incense or medicinal vapor. Powdered bark and other components were used in solutions to treat colds. Resin was used to dress wounds or chewed as gum. The tree boughs were used for bedding. Some
Plateau Indian Indigenous peoples of the Northwest Plateau, also referred to by the phrase Indigenous peoples of the Plateau, and historically called the Plateau Indians (though comprising many groups) are Indigenous peoples of the Americas, indigenous peoples ...
tribes drank or washed in a subalpine fir
boil A boil, also called a furuncle, is a deep folliculitis, which is an infection of the hair follicle. It is most commonly caused by infection by the bacterium '' Staphylococcus aureus'', resulting in a painful swollen area on the skin caused by ...
for purification or to make their hair grow. The light wood is considered poor quality, but sometimes used for wood pulp, general structural purposes and paper manufacture. It is also a popular
Christmas tree A Christmas tree is a decorated tree, usually an evergreen conifer, such as a spruce, pine or fir, or an artificial tree of similar appearance, associated with the celebration of Christmas. The custom was further developed in early modern ...
. It is a popular
ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
for parks and large gardens, grown for its strongly glaucous-blue foliage. It can also function as a
bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of '' penjing''. Unlike ''penjing'', which utilizes traditional techniques to produc ...
. The
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
''Abies lasiocarpa'' var. ''arizonica'' 'Compacta' is suitable for smaller gardens, growing as a shrub to tall by broad. In the UK It has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
. The largest-known specimen, measuring thick and tall, had a small door in its trunk and a storage space, which the film crew of the
Disney The Walt Disney Company, commonly known as Disney (), is an American multinational mass media and entertainment conglomerate headquartered at the Walt Disney Studios complex in Burbank, California. Disney was originally founded on October ...
-produced documentary ''
The Olympic Elk ''The Olympic Elk'' is a 1952 American short documentary film directed by James Algar and produced by Walt Disney as part of the ''True-Life Adventures'' series of nature documentaries. Summary A photographic study of the Olympic elk which abou ...
'' (1952) used to store equipment. HurricaneRidge 7392t.jpg, Specimen in
Olympic National Park Olympic National Park is a United States national park located in the State of Washington, on the Olympic Peninsula. The park has four regions: the Pacific coastline, alpine areas, the west-side temperate rainforest, and the forests of the drier ...
in mid-September Abies lasiocarpa 7458.jpg, Specimen in
Mount Rainier National Park Mount Rainier National Park is an American national park located in southeast Pierce County and northeast Lewis County in Washington state. The park was established on March 2, 1899, as the fourth national park in the United States, preservin ...
in late September


References


External links

* *
Gymnosperm Database: ''Abies lasiocarpa''Flora of North America: ''Abies lasiocarpa''Flora of North America: ''Abies bifolia''Interactive Distribution Map of ''Abies lasiocarpa''
{{Taxonbar, from=Q978755 lasiocarpa Trees of Western Canada Trees of the Western United States Flora of the Rocky Mountains Trees of Alberta Trees of the Northwestern United States Flora of the Klamath Mountains Taxa named by Thomas Nuttall Trees of North America Trees of the United States Trees of Canada Trees of Alaska