Struthiocephaloides
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''Struthiocephalus'' ("Ostrich Head") is an extinct genus of
dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephal ...
n
therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...
s from the
Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. It was a large animal, reaching in body mass.


Ecology

This animal has the largest head of any tapinocephalid. Brink (1956) suggests that ''Struthiocephalus'' fed in or near water, the teeth being used for rooting up, gathering and grasping plant matter. Boonstra (1965) likewise considered that ''Struthiocephalus'' fed on soft vegetation, possibly under water. He suggested the postcranial skeleton possibly showed adaptations to living in marshy conditions, and the bone surface around the nostril might indicate the presence of a fleshy valve present used for closing off the nostril under water.


Classification

The long-snouted ''Struthiocephalus whaitsi'' would seem to be the only genus and species of this taxon. There are a large number of synonyms. ''Struthiocephalellus'' is apparently a juvenile of ''Struthiocephalus''. Boonstra showed that the seven named species of ''Struthiocephalus'' represent a growth series and are hence synonyms of the first described species, ''S. whaitsi''. ''Struthiocephalus'' is characterized simply by a naso-frontal boss in mature specimens, whilst the very similar genus ''Struthiocephaloides'' lacks this character. This is consistent with a sexually dimorphic character, as well as a role in intraspecific combat. Barghusen (1975) considers would be more effective in flank butting than head-butting per se. In head-on combat the presumed horn would be deflected by the opponent's head and contact would be lost, but in flank butting a horned boss would concentrate the blow. Both ''Struthiocephalus'' and ''Struthiocephaloides'' have the same stratigraphic range - Lower to Middle ''Tapinocephalus'' Zone. Of the remaining two monospecific genera, ''Struthionops intermedius'' known from a single skull, possesses a naso-frontal boss moderate pachyostosis and a fairly short snout. Possibly this is a variant of another species. On the basis of the shorter snout, King (1988) includes this species among the Tapinocephalini. ''Taurocephalus lerouxi'', known from a single skull, has a fairly strong snout and 20 teeth in the upper jaw (more than usual for tapinocephalids). Since tooth count tends to vary greatly among individual dinocephalians, again only this is also likely a variant individual, most probably ''Struthiocephalus''. The remaining species, ''Moschosaurus longiceps'' is known from a single small and lightly built skull about long from the Upper ''Tapinocephalus'' Zone. It was originally placed in its own family, the Moschosauridae, and considered a good ancestral or primitive form (despite its late date). However, Boonstra (1969) identifies it as a juvenile ''Struthiocephalus''.


See also

*
List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ...


References


Further reading

*''Dinosaur Encyclopedia'' by Jayne Parsons *''The Origin and Evolution of Mammals'' (Oxford Biology) by T. S. Kemp *''Reptiles and Herbivory'' by Gillian King *''Origins of the Higher Groups of Tetrapods: Controversy and Consensus'' by Hans-Peter Schultze and Linda Trueb


External links


''Tapinocephalidae''
at Paleos Tapinocephalians Prehistoric therapsid genera Guadalupian synapsids Guadalupian synapsids of Africa Fossil taxa described in 1915 Taxa named by Sidney H. Haughton Capitanian genus first appearances Capitanian genus extinctions {{Paleo-therapsid-stub