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botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, a drupe (or stone fruit) is an
indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
in which an outer fleshy part (
exocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
, or skin, and
mesocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Agg ...
, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened
endocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
with a
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
(''kernel'') inside. These fruits usually develop from a single
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils' ...
, and mostly from flowers with superior ovaries (
polypyrenous drupe In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and re ...
s are exceptions). The definitive characteristic of a drupe is that the hard, lignified stone is derived from the ovary wall of the
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
. In an
aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
, which is composed of small, individual drupes (such as a raspberry), each individual is termed a drupelet, and may together form an aggregate fruit. Such fruits are often termed ''
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, rasp ...
'', although botanists use a different definition of ''berry''. Other fleshy fruits may have a stony enclosure that comes from the seed coat surrounding the seed, but such fruits are not drupes.
Flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s that produce drupes include
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
,
jujube Jujube (), sometimes jujuba, known by the scientific name ''Ziziphus jujuba'' and also called red date, Chinese date, and Chinese jujube, is a species in the genus '' Ziziphus'' in the buckthorn family Rhamnaceae. Description It is a smal ...
, mango, olive, most palms (including açaí,
date Date or dates may refer to: *Date (fruit), the fruit of the date palm (''Phoenix dactylifera'') Social activity *Dating, a form of courtship involving social activity, with the aim of assessing a potential partner ** Group dating *Play date, a ...
,
sabal ''Sabal'' is a genus of palms (or fan-palms) endemic to the New World. Currently, there are 17 recognized species of ''Sabal'', including one hybrid species. The species are native to the subtropical and tropical regions of the Americas, from th ...
, coconut and
oil palm ''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm '' Elaeis guineensis'' (the species name ''guineensis'' referring to its c ...
s), pistachio,
white sapote The white sapote, scientific name ''Casimiroa edulis'', also called casimiroa and Mexican apple, and known as ''cochitzapotl'' in the Nahuatl language (meaning "sleep-sapote") is a species of tropical fruiting tree in the family Rutaceae, nativ ...
, cashew, and all members of the genus ''
Prunus ''Prunus'' is a genus of trees and shrubs, which includes (among many others) the fruits plums, Cherry, cherries, peaches, Peach#Nectarine, nectarines, apricots, and almonds. Native to the North American temperate regions, the neotropics of Sou ...
'', including the almond, apricot, cherry,
damson The damson () or damson plum ('' Prunus domestica'' subsp. ''insititia'', or sometimes ''Prunus insititia''),M. H. Porche"Sorting ''Prunus'' names" in "Multilingual multiscript plant names database, University of Melbourne. Plantnames.unimelb.e ...
,
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-f ...
,
nectarine The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-f ...
, and plum. The term drupaceous is applied to a fruit having the structure and texture of a drupe, but which does not precisely fit the definition of a drupe.


Terminology

The boundary between a drupe and a berry is not always clear. Thus, some sources describe the fruit of species of the genus '' Persea'', which includes the
avocado The avocado (''Persea americana'') is a medium-sized, evergreen tree in the laurel family ( Lauraceae). It is native to the Americas and was first domesticated by Mesoamerican tribes more than 5,000 years ago. Then as now it was prized for ...
, as a drupe, others describe avocado fruit as a berry. One definition of ''berry'' requires the endocarp to be less than thick, other fruits with a stony endocarp being drupes. In marginal cases, terms such as ''drupaceous'' or ''drupe-like'' may be used. The term stone fruit (also stonefruit) can be a synonym for drupe or, more typically, it can mean just the fruit of the genus ''Prunus''. Freestone refers to a drupe having a stone which can be removed from the flesh with ease. The flesh is not attached to the stone and does not need to be cut to free the stone. Freestone varieties of fruits are preferred for uses that require careful removal of the stone, especially if removal will be done by hand. Freestone plums are preferred for making homegrown
prune A prune is a dried plum, most commonly from the European plum (''Prunus domestica''). Not all plum species or varieties can be dried into prunes. A prune is the firm-fleshed fruit (plum) of '' Prunus domestica'' varieties that have a high so ...
s, and freestone sour cherries are preferred for making pies and
cherry soup Sour cherry soup is a slightly sweet soup made with sour cream, sugar and whole fresh sour cherries, and served chilled. Originating in Hungarian cuisine, this soup is a summer delicacy in several European cuisines. The dish has been adopte ...
. Clingstone refers to a drupe having a stone which cannot be easily removed from the flesh. The flesh is attached strongly to the stone and must be cut to free the stone. Clingstone varieties of fruits in the genus ''Prunus'' are preferred as table fruit and for jams, because the flesh of clingstone fruits tends to be more tender and juicy throughout. Tryma is a specialized term for such
nut Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Co ...
-like drupes that are difficult to categorize. Hickory nuts ( ''Carya'') and
walnut A walnut is the edible seed of a drupe of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. Although culinarily considered a "nut" and used as such, it is not a true ...
s (''
Juglans Walnut trees are any species of tree in the plant genus ''Juglans'', the type genus of the family Juglandaceae, the seeds of which are referred to as walnuts. All species are deciduous trees, tall, with pinnate leaves , with 5–25 leaflets ...
'') in the
Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...
family grow within an outer husk; these fruits are technically drupes or drupaceous nuts, thus are not true botanical nuts.W.P. Armstrong. 2008
''Identification Of Major Fruit Types''
/ref>


Ecology

Many drupes, with their sweet, fleshy outer layer, attract the attention of animals as a food, and the plant population benefits from the resulting dispersal of its seeds. The
endocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
(pit or stone) is sometimes dropped after the fleshy part is eaten, but is often swallowed, passing through the
digestive tract The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and ...
, and returned to the soil in feces with the seed inside unharmed. This passage through the digestive tract can reduce the thickness of the endocarp, thus can aid in germination rates. The process is known as
scarification Scarification involves scratching, etching, burning/branding, or superficially cutting designs, pictures, or words into the skin as a permanent body modification or body art. The body modification can take roughly 6–12 months to heal. In the ...
.


Examples

Typical drupes include
apricots An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus '' Prunus''. Usually, an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are al ...
,
olives The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' 'M ...
,
loquat The loquat (''Eriobotrya japonica'') is a large evergreen shrub or tree, grown commercially for its orange fruit and for its leaves, which are used to make herbal tea. It is also cultivated as an ornamental plant. The loquat is in the family R ...
,
peaches The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-fu ...
,
plums A plum is a fruit of some species in ''Prunus'' subg. ''Prunus'.'' Dried plums are called prunes. History Plums may have been one of the first fruits domesticated by humans. Three of the most abundantly cultivated species are not found i ...
, cherries,
mangoes A mango is an edible stone fruit produced by the tropical tree ''Mangifera indica''. It is believed to have originated in the region between northwestern Myanmar, Bangladesh, and northeastern India. ''M. indica'' has been cultivated in South ...
,
pecans The pecan (''Carya illinoinensis'') is a species of hickory native to the southern United States and northern Mexico in the region of the Mississippi River. The tree is cultivated for its seed in the southern United States, primarily in Georgia, ...
, and amlas (Indian gooseberries). Other examples include sloe (''
Prunus spinosa ''Prunus spinosa'', called blackthorn or sloe, is a species of flowering plant in the rose family Rosaceae. The species is native to Europe, western Asia, and regionally in northwest Africa. It is locally naturalized in New Zealand, Tasmania, ...
'') and ivy (''
Hedera helix ''Hedera'', commonly called ivy (plural ivies), is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to western, central and southern Europe, Macaronesia, northwestern Africa and ...
''). The coconut is also a drupe, but the
mesocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Agg ...
is fibrous or dry (termed a ''husk''), so this type of fruit is classified as a simple dry, fibrous drupe. Unlike other drupes, the coconut seed is so large that it is unlikely to be dispersed by being swallowed by
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''Biota (ecology ...
, but it can float extremely long distances—across oceans.
Bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...
fruits such as the
blackberry The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy ...
and the raspberry are aggregates of drupelets. The fruit of blackberries and raspberries comes from a single flower whose pistil is made up of a number of free carpels. However,
mulberries ''Morus'', a genus of flowering plants in the family Moraceae, consists of diverse species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 identif ...
, which closely resemble blackberries, are not aggregate fruit, but are multiple fruits, actually derived from bunches of catkins, each drupelet thus belonging to a different flower. Certain drupes occur in large clusters, as in the case of palm species, where a large array of drupes is found in a cluster. Examples of such large drupe clusters include
dates Date or dates may refer to: *Date (fruit), the fruit of the date palm (''Phoenix dactylifera'') Social activity *Dating, a form of courtship involving social activity, with the aim of assessing a potential partner **Group dating * Play date, a ...
, ''
Jubaea chilensis ''Jubaea'' is a genus of palms with one species, ''Jubaea chilensis'' or ''Jubaea spectabilis'', commonly known in English as the Chilean wine palm or Chile cocopalm, and palma chilena in Spanish. It is native to southwestern South America and i ...
'' in central Chile and ''
Washingtonia filifera ''Washingtonia filifera'', the desert fan palm, California fan palm, or California palm,Flora of North America Association. ''Flora of North America: North of Mexico Volume 22: Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in Part), and Zi ...
'' in the
Sonoran Desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Ariz ...
of North America. Drupe-like "fruits" are also known in many
gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμν ...
s like
cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk with a crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male o ...
s,
ginkgo ''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus with ...
s and some cypresses.


Gallery

Image:NIEdot325.jpg, Assorted drupes Image:Autumn Red peaches.jpg, The
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-f ...
is a typical drupe (stone fruit) Image:Prunus - Elena.JPG, 'Elena', a freestone
prune plum The prune plum (''Prunus domestica'' subsp. ''domestica'') is a fruit-bearing tree, or its fruit. It is a subspecies of the plum ''Prunus domestica''. The freestone fruit is similar to, but distinct from, the clingstone damson (''Prunus domesti ...
Image:Nectarine_stone.jpg, The pit of a
nectarine The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-f ...
Image:Black Pepper (Piper nigrum) fruits.jpg, Unripe drupes of
black pepper Black pepper (''Piper nigrum'') is a flowering vine in the family Piperaceae, cultivated for its fruit, known as a peppercorn, which is usually dried and used as a spice and seasoning. The fruit is a drupe (stonefruit) which is about in dia ...
Image:Black Butte blackberry.jpg, 'Black Butte' blackberry, a bramble fruit of aggregated drupelets Image:arecanut.jpg, A ripe
areca nut ''Areca'' is a genus of 51 species of palms in the family Arecaceae, found in humid tropical forests from the islands of the Philippines, Malaysia and India, across Southeast Asia to Melanesia. The generic name ''Areca'' is derived from a name ...
File:Ginkgo biloba 007.jpg,
Ginkgo ''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus with ...
"fruits", often noted as drupe-like


See also

* Pome (polypyrenous drupe)


References


External links


Identification Of Major Fruit Types


* {{Nuts Edible nuts and seeds Fruit morphology