Stejneger's pit viper
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''Trimeresursus stejnegeri'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of venomous
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
endemic to Asia. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are currently recognized, including the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
described here.
Common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
s for this pit viper include Stejneger's pit viper, Chinese pit viper, Chinese green tree viper,U.S. Navy (1991). ''Poisonous Snakes of the World''. New York: US Government / Dover Publications Inc. 203 pp. . bamboo viper, Chinese bamboo pitviper, 69 bamboo viper, and Chinese tree viper. Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . For other common, non-scientific names, see below.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''stejnegeri'', is in honor of
Leonhard Stejneger Leonhard Hess Stejneger (30 October 1851 – 28 February 1943) was a Norwegian-born American ornithologist, herpetologist and zoologist. Stejneger specialized in vertebrate natural history studies. He gained his greatest reputation with reptiles ...
, the Norwegian-born,
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
herpetologist who worked at the Smithsonian Institution for over 60 years.


Description

''Trimeresurus stejnegeri'' grows to a maximum total length of , which includes a tail length of . The males have
hemipenes A hemipenis (plural hemipenes) is one of a pair of intromittent organs of male squamata, squamates (snakes, lizards and amphisbaenia, worm lizards). Hemipenes are usually held inverted within the body, and are everted for reproduction via erecti ...
that are short and spinose beyond the bifurcation. Leviton AE, Wogan GOU, Koo MS, Zug GR, Lucas RS, Vindum JV (2003). "The Dangerously Venomous Snakes of Myanmar, Illustrated Checklist with Keys". ''Proceedings of the California Academy of Sciences'' 54 (24): 407-462. The dorsal scales are arranged in 21 longitudinal rows at midbody. There are 9–11
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
, of which the first are separated from nasal scales by a distinct suture. The
supraocular In scaled reptiles, supraocular scales are (enlarged) scales on the crown immediately above the eye.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. ...
s are single, narrow, and sometimes divided by a transverse suture. There are 11–16 scales in a line between the supraoculars. The
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
number 150–174, and the subcaudals are 54–77. All of the subcaudals are paired. The color pattern is bright to dark green above, pale green to whitish below, the two separated by a bright bicolored orange or brown (below) and white (above) (males) or bicolored or white only (females) ventrolateral stripe, which occupies the whole of the outermost scale row and a portion of the second row. Bamboo vipers are carnivores: they eat small rodents, birds, frogs, and lizards. Yellow colored mutants have been reported.


Common names

Common names for ''T. stejnegeri'' include bamboo viper, Chinese tree viper, bamboo snake, Chinese green tree viper, Chinese bamboo viper, Stejneger's pit viper, Stejneger's palm viper, red tail snake, Stejneger's bamboo pitviper, Gumprecht A, Tillack F, Orlov NL,
Captain A Captain is a title, an appellative for the commanding officer of a military unit; the supreme leader of a navy ship, merchant ship, aeroplane, spacecraft, or other vessel; or the commander of a port, fire or police department, election precinct, e ...
, Ryabov S (2004). ''Asian Pitvipers''. First Edition. Berlin: Geitje Books. Berlin. 368 pp. .


Geographic range

''Trimeresurus stejnegeri'' is found in Northeast India) and Nepal through
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
and
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
to much of southern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
( Yunnan, Sichuan,
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...
, Jiangxi, Jiangsu, Hunan, Hubei, Guizhou,
Guangxi Guangxi (; ; Chinese postal romanization, alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an Autonomous regions of China, autonomous region of the People's Republic ...
, Guangdong, Hainan, Fujian, Anhui, Zhejiang), Vietnam, and Taiwan. The type locality was originally listed as "Shaowu, Fukien Province, China", and later emended to "N.W. Fukien Province" by Pope & Pope (1933) (Fukien being the former romanization of Fujian).


Habitat

The preferred natural habitat of ''T. stejnegeri'' is forest, at altitudes from sea level to .


Venom

''Trimeresurus stejnegeri'' has a potent hemotoxin. The wound usually feels extremely painful, as if it had been branded with a hot iron, and the pain does not subside until about 24 hours after being bitten. Within a few minutes of being bitten, the surrounding flesh dies and turns black, highlighting the puncture wounds. The wound site quickly swells, and the skin and muscle become black due to
necrosis Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. Necrosis is caused by factors external to the cell or tissue, such as infection, or trauma which result in the unregulated dige ...
. The size of the necrotic area depends on the amount of venom injected and the depth of the bite.


Reproduction

''T. stejnegeri'' is viviparous.


Subspecies


References


Further reading

*Creer S, Malhotra A, Thorpe RS, Chou WH (2001). "Multiple causation of phylogeographical pattern as revealed by nested clade analysis of the bamboo viper (''Trimeresurus stejnegeri'') within Taiwan". ''Molecular Ecology'' 10 (8): 1967-1981. * Das I (2002). ''A Photographic Guide to Snakes and other Reptiles of India''. Sanibel Island, Florida: Ralph Curtis Books. 144 pp. . (''Trimeresurus stejnegeri'', p. 68). *Malhotra A, Thorpe RS (2004). "Maximizing information in systematic revisions: a combined molecular and morphological analysis of a cryptic green Pit Viper complex (''Trimeresurus stejnegeri'')". ''Biological Journal of the Linnean Society'' 82 (2): 219. * Parkinson CL (1999). "Molecular systematics and biogeographical history of Pit Vipers as determined by mitochondrial ribosomal DNA sequences". ''Copeia'' 1999 (3): 576–586. * Guo P, Zhang F (2001). "Comparative studies on hemipenes of four species of ''Trimeresurus'' (''sensu stricto'') (Serpentes: Crotalinae)". ''Amphibia-Reptilia'' 22 (1): 113-117. * Schmidt KP (1925). "New Reptiles and a New Salamander from China". ''American Museum Novitates'' (157): 1-5. ("''Trimeresurus stejnegeri'', new species", p. 4). *Tu M-C et al. (2000). "Phylogeny, Taxonomy, and Biogeography of the Oriental Pit Vipers of the Genus ''Trimeresurus'' (Reptilia: Viperidae: Crotalinae): A Molecular Perspective". ''Zoological Science'' 17: 1147–1157. {{Taxonbar, from=Q2719425 stejnegeri Snakes of Asia Reptiles of Cambodia Snakes of China Reptiles of India Reptiles of Laos Reptiles of Myanmar Reptiles of Taiwan Snakes of Vietnam Taxa named by Karl Patterson Schmidt Reptiles described in 1925