Steenbok
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The steenbok (''Raphicerus campestris'') is a common small antelope of southern and eastern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. It is sometimes known as the steinbuck or steinbok.


Description

Steenbok resemble small
oribi The oribi (; ''Ourebia ourebi'') is a small antelope found in eastern, southern and western Africa. The sole member of its genus, it was described by the German zoologist Eberhard August Wilhelm von Zimmermann in 1783. While this is the only ...
, standing 45–60 cm (16"–24") at the shoulder, and average ~12 kg. Their coat is any shade from fawn to rufous, typically rather orange. The underside, including chin and throat, is white, as is the ring around the eye. Ears are large with "finger-marks" on the inside. Males have straight, smooth, parallel
horns Horns or The Horns may refer to: * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * ''Horns'' (novel), a dark fantasy novel written in 2010 by Joe Hill ...
7–19 cm long (see image left). There is a black crescent-shape between the ears, a long black bridge to the glossy black nose, and a black circular scent-gland in front of the eye. The tail is not usually visible, being only 4–6 cm long.


Distribution

There are two distinct clusters in steenbok distribution. In East Africa, it occurs in central and southern
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
and northern
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
. It was formerly widespread in
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
, but is now almost certainly extinct there. In
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
, it occurs in
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
,
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
, Eswatini,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label= Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalaha ...
,
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...
,
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and ...
and probably Lesotho.


Habitat

Steenbok live in a variety of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s from semi-desert, such as the edge of the
Kalahari Desert The Kalahari Desert is a large semi-arid sandy savanna in Southern Africa extending for , covering much of Botswana, and parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal d ...
and
Etosha National Park Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
, to open woodland and thickets, including open
plain In geography, a plain is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and as plateaus or uplands ...
s, stony
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
h, and '' Acacia''–
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
mosaic A mosaic is a pattern or image made of small regular or irregular pieces of colored stone, glass or ceramic, held in place by plaster/mortar, and covering a surface. Mosaics are often used as floor and wall decoration, and were particularly pop ...
s. They are said to favour unstable or transitional habitats.Kingdon, Jonathan. 1997. ''The Kingdon Field Guide to African Mammals''. Academic Press, San Diego & London. Pp. 387–388. () At least in the central part of Kruger National Park, South Africa, Steenbok show a distinct preference for ''
Acacia tortilis ''Vachellia tortilis'', widely known as ''Acacia tortilis'' but now attributed to the genus '' Vachellia'', is the umbrella thorn acacia, also known as ''umbrella thorn'' and Israeli babool, a medium to large canopied tree native to most of Afr ...
'' savannah throughout the year, with no tendency to migrate to moister areas during the dry season (unlike many larger African savannah ungulates, including species sympatric with Steenbok in the wet season). Population density is typically 0.3–1.0 individuals per square kilometre, reaching 4 per km2 in optimal habitats.Kleiman, David G. ''et al.'', Eds. 2003. ''Grzimek's Animal Life Encyclopedia'', 2nd edn. Vol. 16: Mammals V. Gale Cengage Learning. Pp. 59–72.


Diet

Steenbok typically
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
on low-level vegetation (they cannot reach above 0.9 m), but are also adept at scraping up roots and tubers. In central Kruger National Park, Steenbok show a distinct preference for
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s, and then woody plants (especially ''
Flueggea virosa ''Flueggea'', the bushweeds, is a genus of shrubs and trees in the family Phyllanthaceae first described as a genus in 1806. It is widespread across much of Asia, Africa, and various oceanic islands, with a few species in South America and on th ...
'') when few forbs are available. They will also take fruits and only very rarely graze on grass. They are almost entirely independent of drinking water, gaining the moisture they need from their food.


Behaviour

Steenbok are active during the day and the night; however, during hotter periods, they rest under shade during the heat of the day. The time spent feeding at night increases in the dry season. While resting, they may be busy grooming,
ruminating Ruminants (suborder Ruminantia) are ungulate, hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principally ...
or taking brief spells of sleep.


Anti-predator

At the first sign of trouble, steenbok typically lie low in the vegetation. If a predator or perceived threat comes closer, a steenbok will leap away and follow a zigzag route to try to shake off the pursuer. Escaping steenbok frequently stop to look back, and flight is alternated with prostration during extended pursuit. They are known to take refuge in the burrows of
aardvarks The aardvark ( ; ''Orycteropus afer'') is a medium-sized, burrowing, nocturnal mammal native to Africa. It is the only living species of the order Tubulidentata, although other prehistoric species and genera of Tubulidentata are known. Unlik ...
. Known predators include Southern African wildcat,
caracal The caracal (''Caracal caracal'') () is a medium-sized wild cat native to Africa, the Middle East, Central Asia, and arid areas of Pakistan and northwestern India. It is characterised by a robust build, long legs, a short face, long tufted e ...
,
jackals Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed ...
, leopard,
martial eagle The martial eagle (''Polemaetus bellicosus'') is a large eagle native to sub-Saharan Africa.Ferguson-Lees & Christie, ''Raptors of the World''. Houghton Mifflin Company (2001), . It is the only member of the genus ''Polemaetus''. A species of t ...
and
pythons The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 42 species are currently recognized. Distribution ...
.


Breeding

Steenbok are typically solitary, except for when a pair come together to mate. However, it has been suggested that pairs occupy consistent
territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
while living independently, staying in contact through
scent marking In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
s, so that they know where their mate is most of the time. Scent marking is primarily through
dung midden Dung middens, also known as ''dung hills'',''The New Encyclopaedia of Mammals'' D MacDonald 2002 Oxford are piles of dung that mammals periodically return to and build up. They are used as a form of territorial marker. A range of animals are know ...
s. Territories range from 4 hectares to 1 square kilometre. The male is aggressive during the female's
oestrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...
, engaging in "bluff-and-bluster" type displays with rival males—prolonged contests invariably involve well-matched individuals, usually in their prime.Cohen, Michael. 1976. The Steenbok: A neglected species. ''Custos'' (April 1976): 23–26. Breeding occurs throughout the year, although more fawns are born November to December in the southern spring–summer; some females may breed twice a year. Gestation period is about 170 days, and usually a single
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
fawn is produced. The fawn is kept hidden in vegetation for 2 weeks, but suckles for 3 months. Females become sexually mature at 6–8 months and males at 9 months. Steenbok are known to live for 7 years or more.


Taxonomy

Two subspecies are recognized: ''R. c. campestris'' in Southern Africa and ''R. c. neumanni'' of East Africa; although
MSW3 ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and bibliographic data for the known species of mammals. It is now in its third edition, published in late 2005 ...
also recognizes ''capricornis'' and ''kelleni''. Up to 24 subspecies have been described from Southern Africa, distinguished on such features as coat colour.


Gallery

Steenbok female.jpg , Female in South Africa Steenbok (Raphicerus campestris) male.jpg, Backlit male showing white fur in ears, Tswalu Kalahari Res. Steenbok (Raphicerus campestris) running composite.jpg, Female running in Damaraland,
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...


Notes


References


External links


Wild About You (info and image)


{{Taxonbar, from=Q368033 Dwarf antelopes Fauna of East Africa Mammals of Angola Mammals of Botswana Mammals of Kenya Mammals of Mozambique Mammals of Namibia Mammals of South Africa Mammals of Eswatini Mammals of Tanzania Mammals of Zambia Mammals described in 1811