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Staphyliniformia
Staphyliniformia
is a large infraorder of beetles. It contains over 60,000 described species from all regions of the world. Most species occur in moist habitats - various kinds of rotting plant debris, fungi, dung, carrion, many live in fresh water.

Contents

1 Characteristics 2 Systematics and evolution 3 See also 4 References

Characteristics[edit] Most Staphyliniforms are small to average sized beetles. The diverse group has few clear apomorphies. They have primitively 11-segmented antennae, a constricted neck well behind the eyes. The pronotum has a well defined, large lateral edge. Larval legs are 5-segmented, the 10th abdominal segment is often with more-or-less fine or strong spines or hooks. Urogomphi (paired "horns" at posterior tip of abdomen of larvae and pupae) with basal articulation.[1] Systematics and evolution[edit] Staphyliniformia
Staphyliniformia
belongs to the Polyphaga
Polyphaga
suborder and is usually given an infraorder or series rank. It contains three superfamilies:

Histeroidea, including the clown beetles. Hydrophiloidea, including the water scavenger beetles Staphylinoidea, including antlike stone beetles, carrion beetles, and rove beetles.

Some recent studies also include the superfamily Scarabaeoidea (infraorder Scarabaeiformia), forming together the so-called Hydrophiloid lineage.[2][3] A sister group relationship of Hydrophiloidea
Hydrophiloidea
and Histeroidea
Histeroidea
is strongly supported.[4] The unambiguous fossil record dates back to Triassic, and an early Mesozoic
Mesozoic
origin of the group is probable.[5] See also[edit] List of subgroups of the order Coleoptera References[edit]

^ M.Hansen. Phylogeny and classification of the staphyliniform beetle families (Coleoptera). Biologiske Skrifter 48, Copenhagen, 1997 ^ M.S.Caterino et al., On the constitution and phylogeny of Staphyliniformia
Staphyliniformia
(Insecta: Coleoptera). Molecular Phylogenetics and Evolution Vol.34, 3, 2005 ^ J.Kukalova-Peck, J.F.Lawrence, (1993) Evolution of the hind wings in Coleoptera. Canadian Entomologist, 135, 181–258. ^ R.G.Beutel, R.A.B.Leschen, Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology (2005), 30, 510–548 ^ Grimaldi, D. and Engel, M.S. (2005). Evolution of the Insects. Cambridge University Press

Data related to Staphyliniformia
Staphyliniformia
at Wikispecies

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Extant Coleoptera families

Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera Superorder: Endopterygota

Suborder Archostemata

Crowsoniellidae (Crowsoniella relicta) Cupedidae
Cupedidae
(reticulated beetles) Jurodidae (Sikhotealinia zhiltzovae) Micromalthidae (telephone-pole beetle) Ommatidae

Suborder Adephaga

Extant families

Amphizoidae (trout-stream beetles) Aspidytidae Carabidae (ground beetles) Dytiscidae
Dytiscidae
(predaceous diving beetles) Gyrinidae (whirligig beetles) Haliplidae
Haliplidae
(crawling water beetles) Hygrobiidae Meruidae (Meru phyllisae) Noteridae
Noteridae
(burrowing water beetles) Rhysodidae
Rhysodidae
(wrinkled bark beetles) Trachypachidae
Trachypachidae
(false ground beetles)

Suborder Myxophaga

Hydroscaphidae
Hydroscaphidae
(skiff beetles) Lepiceridae Sphaeriusidae Torridincolidae

Suborder Polyphaga

Bostrichiformia

Bostrichoidea

Anobiidae
Anobiidae
(furniture beetles, death watch beetles, spider beetles) Bostrichidae
Bostrichidae
(auger beetles) Dermestidae
Dermestidae
(skin beetles) Jacobsoniidae
Jacobsoniidae
(Jacobson's beetles) Nosodendridae
Nosodendridae
(wounded-tree beetles)

Derodontoidea

Derodontidae
Derodontidae
(tooth-necked fungus beetles)

Cucujiformia

Chrysomeloidea

Cerambycidae (longhorn beetles) Chrysomelidae (leaf beetles) Disteniidae Megalopodidae Orsodacnidae Oxypeltidae Vesperidae

Cleroidea

Acanthocnemidae (Acanthocnemus nigricans) Chaetosomatidae Cleridae
Cleridae
(checkered beetles) Melyridae
Melyridae
(soft-wing flower beetles) Phloiophilidae (Phloiophilus edwardsi) Phycosecidae Prionoceridae Trogossitidae
Trogossitidae
(bark-gnawing beetles)

Cucujoidea

Alexiidae Biphyllidae
Biphyllidae
(false skin beetles) Boganiidae Bothrideridae
Bothrideridae
(dry bark beetles) Byturidae
Byturidae
(fruitworm beetles) Cavognathidae Cerylonidae
Cerylonidae
(minute bark beetles) Coccinellidae
Coccinellidae
(lady beetles, or God's cows) Corylophidae
Corylophidae
(minute fungus beetles) Cryptophagidae
Cryptophagidae
(silken fungus beetles) Cucujidae
Cucujidae
(flat bark beetles) Cyclaxyridae Discolomatidae Endomychidae
Endomychidae
(handsome fungus beetles) Erotylidae
Erotylidae
(pleasing fungus beetles) Helotidae Hobartiidae Kateretidae
Kateretidae
(short-winged flower beetles) Laemophloeidae
Laemophloeidae
(lined flat bark beetles) Lamingtoniidae (Lamingtonium binnaberrense) Latridiidae
Latridiidae
(minute brown scavenger beetles) Monotomidae (root-eating beetles) Myraboliidae Nitidulidae (sap beetles) Passandridae
Passandridae
(parasitic flat bark beetles) Phalacridae
Phalacridae
(shining flower beetles) Phloeostichidae Propalticidae Protocucujidae Silvanidae
Silvanidae
(silvanid flat bark beetles) Smicripidae (palmetto beetles) Sphindidae
Sphindidae
(dry-fungus beetles)

Curculionoidea (weevils)

Anthribidae
Anthribidae
(fungus weevils) Attelabidae
Attelabidae
(leaf-rolling weevils) Belidae
Belidae
(primitive weevils) Brentidae
Brentidae
(straight snout weevils, New York weevil) Caridae Curculionidae
Curculionidae
(true weevils, bark beetles, ambrosia beetles) Nemonychidae
Nemonychidae
(pine flower weevils)

Lymexyloidea

Lymexylidae
Lymexylidae
(ship-timber beetles)

Tenebrionoidea

Aderidae
Aderidae
(ant-like leaf beetles) Anthicidae
Anthicidae
(ant-like flower beetles) Archeocrypticidae (cryptic fungus beetles) Boridae
Boridae
(conifer bark beetles) Chalcodryidae Ciidae
Ciidae
(minute tree-fungus beetles) Melandryidae
Melandryidae
(false darkling beetles) Meloidae (blister beetles) Mordellidae
Mordellidae
(tumbling flower beetles) Mycetophagidae
Mycetophagidae
(hairy fungus beetles) Mycteridae (palm and flower beetles) Oedemeridae
Oedemeridae
(false blister beetle) Perimylopidae, or Promecheilidae Prostomidae
Prostomidae
(jugular-horned beetles) Pterogeniidae Pyrochroidae (fire-coloured beetles) Pythidae
Pythidae
(dead log bark beetles) Ripiphoridae
Ripiphoridae
(wedge-shaped beetles) Salpingidae
Salpingidae
(narrow-waisted bark beetles) Scraptiidae
Scraptiidae
(false flower beetles) Stenotrachelidae
Stenotrachelidae
(false longhorn beetles) Synchroidae
Synchroidae
(synchroa bark beetles) Tenebrionidae (darkling beetles) Tetratomidae (polypore fungus beetles) Trachelostenidae Trictenotomidae Ulodidae Zopheridae
Zopheridae
(ironclad beetles, cylindrical bark beetles)

Elateriformia

Buprestoidea

Buprestidae
Buprestidae
(jewel beetles, or metallic wood-boring beetles) Schizopodidae

Byrrhoidea

Byrrhidae
Byrrhidae
(pill beetles) Callirhipidae (cedar beetles) Chelonariidae (turtle beetles) Cneoglossidae Dryopidae
Dryopidae
(long-toed water beetles) Elmidae
Elmidae
(riffle beetles) Eulichadidae (forest stream beetles) Heteroceridae
Heteroceridae
(variegated mud-loving beetles) Limnichidae (minute mud beetles) Lutrochidae (travertine beetles) Psephenidae (water-penny beetles) Ptilodactylidae

Dascilloidea

Dascillidae (soft bodied plant beetles) Rhipiceridae
Rhipiceridae
(cicada beetle, cicada parasite beetles)

Elateroidea

Anischiidae Artematopodidae
Artematopodidae
(soft-bodied plant beetles) Brachypsectridae (Texas beetles) Cantharidae (soldier beetles) Cerophytidae (rare click beetles) Drilidae Elateridae (click beetles) Eucnemidae
Eucnemidae
(false click beetles) Lampyridae (fireflies) Lycidae
Lycidae
(net-winged beetles) Omalisidae Omethidae
Omethidae
(false fireflies) Phengodidae
Phengodidae
(glowworm beetles, long-lipped beetles) Plastoceridae (Plastocerus angulosus) Podabrocephalidae Rhagophthalmidae Rhinorhipidae (Rhinorhipus tamborinensis) Throscidae (false metallic wood-boring beetles)

Scirtoidea

Clambidae Decliniidae (Declinia relicta) Eucinetidae
Eucinetidae
(plate-thigh beetles) Scirtidae

Scarabaeiformia

Scarabaeoidea

Belohinidae ( Belohina inexpectata) Bolboceratidae Ceratocanthidae Diphyllostomatidae (false stag beetles) Geotrupidae
Geotrupidae
(dor beetles) Glaphyridae
Glaphyridae
(bumble bee scarab beetles) Glaresidae (enigmatic scarab beetles) Hybosoridae
Hybosoridae
(scavenger scarab beetles) Lucanidae (stag beetles) Ochodaeidae
Ochodaeidae
(sand-loving scarab beetles) Passalidae
Passalidae
(betsy beetles) Pleocomidae (rain beetles) Scarabaeidae
Scarabaeidae
(scarabs) Trogidae
Trogidae
(hide beetles)

Staphyliniformia

Histeroidea

Histeridae
Histeridae
(clown beetles) Sphaeritidae (false clown beetles) Synteliidae

Hydrophiloidea

Epimetopidae Georissidae (minute mud-loving beetles) Helophoridae Hydrochidae Hydrophilidae
Hydrophilidae
(water scavenger beetles) Spercheidae

Staphylinoidea

Agyrtidae
Agyrtidae
(primitive carrion beetles) Hydraenidae Leiodidae
Leiodidae
(round fungus beetles) Ptiliidae
Ptiliidae
(feather-winged beetles) Scydmaenidae
Scydmaenidae
(ant-like stone beetles) Silphidae
Silphidae
(carrion beetles) Staphylinidae (rove beetles)

List of subgroups of the order Coleoptera

Taxon identifiers

Wd: Q383230 EoL: 2650706 Fauna Europaea: 11098 ITIS: 678301 NCBI: 41085

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