Staphylea
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''Staphylea'', called bladdernuts, is a small genus of 10 or 11 species of flowering plants in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Staphyleaceae Staphyleaceae is a small family of flowering plants in the order Crossosomatales, native to Europe, temperate and tropical Asia and the Americas. The largest genus ''Staphylea'', which gives the family its name, contains the "bladdernut" trees. ...
, native to
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
regions of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. The highest species diversity is in China, where four species occur. They are large
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s, occasionally small trees, growing to 2–5 m tall. The leaves are
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
, arranged in opposite pairs, and
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
, usually with three leaflets, but ''S. pinnata'' and ''S. colchica''. The flowers are produced in drooping terminal
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s 5–10 cm long, with 5–15 flowers on each panicle; the individual flowers are about 1 cm long, with the five
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s and
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s similar in size and in their white or pale pink colour. The fruit is an inflated papery two- or three-lobed capsule 3–10 cm long, containing a few small nut-like seeds.


Species

''
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
'' currently includes: * '' Staphylea affinis'' (Merr. & L.M.Perry) Byng & Christenh. * '' Staphylea arguta'' (Seem.) Byng & Christenh. * '' Staphylea bolanderi'' A.Gray – Sierra bladdernut. Western North America. * '' Staphylea bumalda'' DC. – Japanese bladdernut. Eastern Asia. * '' Staphylea campanulata'' J.Wen * '' Staphylea cochinchinensis'' (Lour.) Byng & Christenh. - Nepal to S. China and Indo-China. * ''
Staphylea colchica ''Staphylea colchica'', the Caucasian bladdernut, Colchis bladdernut, or ka, ჯონჯოლი, or , is an ornamental shrub in the Staphyleaceae family. It is native to western Georgia. Its binomial name is derived from the western Georgi ...
'' Steven – Colchis bladdernut, jonjoli. Eastern Europe. * '' Staphylea emodi'' Wall. ex Brandis – Himalayan bladdernut. Himalaya. * '' Staphylea formosana'' (Nakai) Byng & Christenh. * '' Staphylea forrestii'' Balf.f. – China. * '' Staphylea holocarpa'' Hemsl. – China. * '' Staphylea indochinensis'' (Merr.) Byng & Christenh. - S. China, N. Vietnam * '' Staphylea insignis'' (Tul.) Byng & Christenh. * '' Staphylea japonica'' (Thunb.) Mabb. * '' Staphylea macrosperma'' (C.C.Huang) Byng & Christenh. * '' Staphylea megaphylla'' (Tul.) Byng & Christenh. * ''
Staphylea pinnata ''Staphylea pinnata'', the European bladdernut, is a species of bladdernut native to Europe and naturalized in Britain. Description It is a deciduous shrub growing up to . The species name ''pinnata'' refers to the pinnate leaves. Small, white, ...
'' L. – European bladdernut. Europe. * '' Staphylea pringlei'' S.Watson – Mexico. * '' Staphylea shweliensis'' W.W.Sm. – China. * '' Staphylea subsessilifolia'' (C.Y.Wu) Byng & Christenh. * '' Staphylea ternata'' (Nakai) Byng & Christenh. * '' Staphylea tricornuta'' (Lundell) Byng & Christenh. * '' Staphylea trifolia'' L. – American bladdernut. Eastern North America. * '' Staphylea yuanjiangensis'' K.M.Feng & T.Z.Hsu Some
botanists This is a list of botanists who have Wikipedia articles, in alphabetical order by surname. The List of botanists by author abbreviation is mostly a list of plant taxonomists because an author receives a standard abbreviation only when that auth ...
previously included the closely related
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Turpinia ''Turpinia'' is a genus of trees and shrubs in family Staphyleaceae, native to Asia and North, Central, and South America. Species include: * '' Turpinia affinis'' * '' Turpinia arguta'' * '' Turpinia cochinchinensis'' * '' Turpinia formosana'' ...
'' in ''Staphylea''.


Cultivation and uses

Several species are grown as
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
s for their flowers and peculiar bladder-like fruit. The popular ''Staphylea × elegans'' is a
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
of unknown origin, probably between ''S. colchica'' and ''S. pinnata''. The pickled flowers can be served as a side dish in
Georgian cuisine Georgian cuisine () consists of cooking traditions, techniques, and practices of Georgia. Georgian cuisine has a distinct character, while bearing some similarities with various national cuisines of the Caucasus and the wider Eastern Europe. E ...
. Image:Staphylea-pinnata-flowers.JPG, ''Staphylea pinnata'' flowers Image:Staphylea-pinnata-fruits.JPG, ''Staphylea pinnata'' fruit File:Staphylea pinnata seeds.jpg , ''Staphylea pinnata'' fruit and seeds File:Jonjoli.jpg, ''Staphylea colchica'' pickled flowers


References


External links


EFloras: search results for ''Staphylea''Flora Europaea: ''Staphylea''
- at desert-tropicals.com

{{Taxonbar, from=Q158845 Rosid genera Staphyleaceae