''Spondylosoma'' (meaning "vertebra body") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

avemetatarsalia Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. ...

n archosaur belonging to the clade

Aphanosauria Aphanosauria ("hidden lizards") is an extinct group of reptiles distantly related to dinosaurs (including birds). They are at the base of a group known as Avemetatarsalia, one of two main branches of archosaurs. The other main branch, Pseudosuc ...

from the late

Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic ...

-age

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...

Lower Santa Maria Formation in Paleorrota Geopark,

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

History

Friedrich von Huene based the genus on a fragmentary postcranial skeleton held at the

University of Tübingen The University of Tübingen, officially the Eberhard Karl University of Tübingen (german: Eberhard Karls Universität Tübingen; la, Universitas Eberhardina Carolina), is a public research university located in the city of Tübingen, Baden-W� ...

. This skeleton includes two

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, t ...

, two cervical

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...

e, four dorsal vertebrae, three

sacral Sacral may refer to: *Sacred Sacred describes something that is dedicated or set apart for the service or worship of a deity; is considered worthy of spiritual respect or devotion; or inspires awe or reverence among believers. The property ...

vertebrae,

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eith ...

e, part of a humerus, part of a

femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...

, and part of a pubis. At the time, he thought it was a

prosauropod Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had l ...

.von Huene, F. (1942). ''Die fossilen Reptilien des südamerikanischen Gondwanalandes.'' C.H. Beck:Munich, 342 p.

erman Erman Rašiti may refer to: Given name * Erman Bulucu (born 1989), Turkish footballer * Erman Eltemur (born 1993), Turkish karateka * Erman Güraçar (born 1974), Turkish footballer * Erman Kılıç (born 1983), Turkish footballer * Erman Kunter (b ...

/ref>

With the discovery of the basal dinosaur '' Staurikosaurus'', ''Spondylosoma'' drew attention as a possible relative. Authors went back and forth on the question, considering it either as a basal dinosaur, or as a "

thecodont Thecodontia (meaning 'socket-teeth'), now considered an obsolete taxonomic grouping, was formerly used to describe a diverse "order" of early archosaurian reptiles that first appeared in the latest Permian period and flourished until the end of t ...

" or other basal archosaur. In 2000,

Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosa ...

noted that it lacks dinosaurian characteristics and was probably a

rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...

n more closely related to

rauisuchids Rauisuchidae is a group of large (up to or more) predatory Triassic archosaurs. There is some disagreement over which genera should be included in Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed wheth ...

,Galton, P.M. (2000). Are ''Spondylosoma'' and ''Staurikosaurus'' (Santa Maria Formation, Middle-Upper Triassic, Brasil) the oldest saurischian dinosaurs? ''Paläontologische Zeitschrift'' 74(3):393-423. whereas in 2004 Max Langer disputed this and included ''Spondylosoma'' as a possible basal dinosaur similar to the

herrerasaur Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from dracohors, separate from dinosauria altogether. They are among the oldest known dinosaurs, ...

s (but did not firmly rule out rauisuchian affinities).Langer, M.C. (2004). Basal Saurischia. In: Weishampel, D.B., Dodson, P., and Osmólska, H. (eds.). ''The Dinosauria'' (second edition). University of California Press:Berkeley, 25-46. In his 2009 thesis on archosaur evolution,

Sterling Nesbitt Sterling Nesbitt (born March 25, 1982, in Mesa, Arizona) is an American paleontologist best known for his work on the origin and early evolutionary patterns of archosaurs. He is currently an associate professor at Virginia Tech in the Department of ...

placed Spondylosoma at Archosauria ''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'', noting that the characters used by Galton to tentatively place the genus in Rauisuchidae are also found in Aetosauria, and that the holotype lacks characters to place it in either

Pseudosuchia Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to ...

Ornithodira Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...

. The redescription of '' Teleocrater''Nesbitt et al. 2017. The earliest bird-line archosaurs and the assembly of the dinosaur body plan. Nature. revealed numerous similarities between ''Spondylosoma'' and a few other Triassic taxa leading to their referral to a new clade of archosaurs, Aphanosauria, which is the sister to Ornithodira within Avemetatarsalia.

References

{{Taxonbar, from=Q549748 Middle Triassic archosaurs Middle Triassic reptiles of South America Ladinian genera Triassic Brazil Fossils of Brazil Santa Maria Formation Paleontology in Rio Grande do Sul Fossil taxa described in 1942 Prehistoric avemetatarsalians