Skua
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The skuas are a group of predatory
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s with seven species forming the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Stercorarius'', the only genus in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Stercorariidae. The three smaller skuas, the long-tailed skua, the Arctic skua, and the pomarine skua are called jaegers in
North American English North American English (NAmE, NAE) is the most generalized variety of the English language as spoken in the United States and Canada. Because of their related histories and cultures, plus the similarities between the pronunciations (accents), v ...
. The English word "skua" comes from the Faroese name for the great skua, , with the island of Skúvoy renowned for its colony of that bird. The general Faroese term for skuas is . The word "jaeger" is derived from the German word , meaning "hunter". The genus name ''Stercorarius'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "of dung"; because the food disgorged by other birds when pursued by skuas was once thought to be excrement. Skuas nest on the ground in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
and
Arctic region The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenland), Finland, Iceland, N ...
s, and are long-distance migrants. They have even been sighted at the
South Pole The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is one of the two points where Earth's axis of rotation intersects its surface. It is the southernmost point on Earth and lies antipod ...
.


Biology and habits

Outside the breeding season, skuas take fish,
offal Offal (), also called variety meats, pluck or organ meats, is the organs of a butchered animal. The word does not refer to a particular list of edible organs, which varies by culture and region, but usually excludes muscle. Offal may also refe ...
, and carrion. Many practice
kleptoparasitism Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when f ...
, which comprises up to 95% of the feeding methods of wintering skuas, by chasing gulls, terns and other seabirds to steal their catches, regardless of the size of the species attacked (up to three times heavier than the attacking skua). The larger species, such as the great skua, also regularly kill and eat adult birds, such as puffins and
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s, and have been seen killing birds as large as a grey heron. On the breeding grounds, the three, more slender northern breeding species commonly eat
lemming A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also includ ...
s. Those species that breed in the southern oceans largely feed on fish that can be caught near their colonies. The eggs and chicks of other seabirds, primarily penguins, are an important food source for most skua species during the nesting season. In the southern oceans and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
region, some skua species (especially the
south polar skua The south polar skua (''Stercorarius maccormicki'') is a large seabird in the skua family, Stercorariidae. An older name for the bird is MacCormick's skua, after explorer and naval surgeon Robert McCormick, who first collected the type specimen. ...
) will readily scavenge carcasses at breeding colonies of both penguins and
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammal, marine mammals. They comprise the extant taxon, extant family (biology ...
s. Skuas will also kill live penguin chicks. In these areas, the skuas will often forfeit their catches to the considerably larger and very aggressive
giant petrel Giant petrels form a genus, ''Macronectes'', from the family Procellariidae, which consists of two species. They are the largest birds of this family. Both species are restricted to the Southern Hemisphere, and though their distributions overla ...
s. Skuas have also been observed to directly pilfer
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digest solid food. Immune factors and immune-modula ...
from the
elephant seal Elephant seals are very large, oceangoing earless seals in the genus ''Mirounga''. Both species, the northern elephant seal (''M. angustirostris'') and the southern elephant seal (''M. leonina''), were hunted to the brink of extinction for oi ...
's teat. They are medium to large
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, typically with grey or brown plumage, often with white markings on the wings. The skuas range in size from the long-tailed skua, ''Stercorarius longicauda'', at , to the
brown skua The brown skua (''Stercorarius antarcticus''), also known as the Antarctic skua, subantarctic skua, southern great skua, southern skua, or hākoakoa (Māori), is a large seabird that breeds in the subantarctic and Antarctic zones and moves furthe ...
, ''Stercorarius antarcticus'', at . On average, a skua is about long, and across the wings. They have longish bills with a hooked tip, and webbed feet with sharp claws. They look like large dark gulls, but have a fleshy
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
above the upper mandible. The skuas are strong, acrobatic fliers. They are generally aggressive in disposition. Potential predators approaching their nests will be quickly attacked by the parent birds, which usually target the heads of intruders – a practice known as ' divebombing'.


Taxonomy

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Stercorarius'' was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the parasitic jaeger (''Stercorarius parasiticus'') as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
. Skuas are related to
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s,
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s, auks, and skimmers. In the three smaller species, all nesting exclusively in the Holarctic, breeding adults have the two central tail feathers obviously elongated, and at least some adults have white on the underparts and pale yellow on the neck. These characteristics are not shared by the larger species, all native to the Southern Hemisphere except for the great skua. Therefore, the skuas are often split into two genera, with only the smaller species retained in ''Stercorarius'', and the large species placed in ''Catharacta''. However, based on
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar wor ...
, behavior, and
feather lice A bird louse is any chewing louse (small, biting insects) of order Phthiraptera which parasitizes warm-blooded animals, especially birds. Bird lice may feed on feathers, skin, or blood. They have no wings, and their biting mouth parts distingui ...
, the overall relationship among the species is best expressed by placing all in a single genus. The pomarine and great skuas' mitochondrial DNA (inherited from the mother) is in fact more closely related to each other than it is to either Arctic or long-tailed skuas, or to the Southern Hemisphere species.Cohen, Baker, Belchschmidt, Dittmann, Furness, Gerwin, Helbig, de Korte, Marshall, Palma, Peter, Ramli, Siebold, Willcox, Wilson and Zink (1997)
Enigmatic phylogeny of skuas.
''Proc. Biol. Sci.'' 264(1379):181–190.
Thus, hybridization must have played a considerable role in the evolution of the diversity of Northern Hemisphere skuas.


Species

The genus contains seven species:


References


External links

*
Skua videos
on the Internet Bird Collection {{Authority control Seabirds Taxa named by Mathurin Jacques Brisson