Sialidae
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Alderflies are
megaloptera Megaloptera is an order of insects. It contains the alderflies, dobsonflies and fishflies, and there are about 300 known species. The order's name comes from Ancient Greek, from ''mega-'' (μέγα-) "large" + ''pteryx'' (πτέρυξ) "win ...
n
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Sialidae. They are closely related to the dobsonflies and fishflies as well as to the prehistoric Euchauliodidae. All living alderflies – about 66
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
all together – are part of the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus Genus ( plural genera ) is a taxonomic rank used in the biological classifica ...
Sialinae, which contains between one and seven extant genera according to different scientists' views.


Description

Sialinae have a body length of less than 25 mm (1 inch), long filamentous antennae, and four large dark wings of which the anterior pair is slightly longer than the posterior. They lack ocelli and their fourth tarsal segment is dilated and deeply bilobed. Dead alderfly larvae are used as bait in fishing.fly fishing entomology alderfy entry
/ref>


Life cycle

The females lay a vast number of eggs on grass stems near water. When the larvae are born they drop into the water or the ground nearby it and make their way into their new aquatic biome. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e are aquatic, active, armed with strong sharp mandibles, and breathe by means of seven pairs of abdominal branchial filaments. When full sized, which takes between one and two years, they leave the water and spend a quiescent
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...
l stage on the land before metamorphosis into the sexually mature insect. Adult alderflies stay near to the water in which they had lived when they were younger. Once in their adult stage, they tend to live only 2 to 3 weeks, which they spend mainly in reproducing.


Classification

In addition to the seven living genera, there are several genera of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
alderflies. Family Sialidae *Genus †'' Dobbertinia'' Handlirsch 1920 Green Series, Germany, Early Jurassic (
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toar ...
) *Genus †'' Sharasialis'' Ponomarenko 2012 Shar-Teeg, Mongolia, Late Jurassic ( Tithonian) *Subfamily Sialinae **Genus '' Austrosialis'' Tillyard,
919 __NOTOC__ Year 919 ( CMXIX) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By Place Byzantine Empire * March 25 – Romanos Lekapenos, admiral (''droungarios'') of the Byz ...
/small> ***Species '' Austrosialis ignicollis'' Tillyard,
919 __NOTOC__ Year 919 ( CMXIX) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By Place Byzantine Empire * March 25 – Romanos Lekapenos, admiral (''droungarios'') of the Byz ...
/small> ***Species '' Austrosialis maxmouldsi'' Theischinger, 1983 **Genus †'' Eosialis'' Nel et al., 2002 ***Species †'' Eosialis dorisi'' Nel et al., 2002 **Genus '' Indosialis'' Lestage, 1927 ***Species '' Indosialis bannaensis'' X.-y. Liu et al., 2006 ***Species '' Indosialis beskonakensis'' Nel, 1988 ***Species '' Indosialis indica'' X.-y. Liu et al., 2008 ***Species '' Indosialis minora'' (Banks, 1920) **Genus '' Leptosialis'' Esben-Petersen, 1920 ***Species '' Leptosialis africana Esben-Petersen, 1920'' **Genus '' Nipponosialis'' Kuwayama, 1962 ***Species '' Nipponosialis amamiensis'' Kuwayama, 1964 ***Species '' Nipponosialis jezoensis'' (Okamoto, 1910) ****Subspecies '' Nipponosialis jezoensis jezoensis'' (Okamoto, 1910) ****Subspecies '' Nipponosialis jezoensis kuwayamai'' Hayashi & Suda, 1995 ***Species '' Nipponosialis kumejimae'' (Okamoto, 1910) **Genus †'' Proindosialis'' Nel, 1988 ***Species †'' Proindosialis cantalensis'' Nel, 1988 **Genus '' Protosialis'' van der Weele, 1909 ***Species '' Protosialis afra'' Navás, (1936) ***Species '' Protosialis americana'' (Rambur, 1842) ***Species '' Protosialis australiensis'' (Tillyard, 919) ***Species ''Protosialis australis'' Navás, 1927 ***Species †''Protosialis baltica'' (Wichard, 1997) ***Species ''Protosialis bifasciata'' (Hagen, 1861) ***Species ''Protosialis bimaculata'' Banks, 1920 ***Species ''Protosialis brasiliensis'' Navás, 1936 ***Species †'' Protosialis casca'' (Engel & Grimaldi, 2007) ***Species '' Protosialis chilensis'' (McLachlan, (1871) ***Species '' Protosialis flammata'' Penny, (1982) ***Species '' Protosialis flavicollis'' (Enderlein, 1910) ***Species '' Protosialis hauseri'' Contreras-Ramos et al., 2005 ***Species '' Protosialis madegassa'' Navás, 1927 ***Species '' Protosialis mexicana'' (Banks, 1901) ***Species '' Protosialis minora'' Banks, 1920 ***Species '' Protosialis nubila'' Navás, 1933 ***Species '' Protosialis ranchograndis'' Contreras-Ramos, 2006 ***Species †'' Protosialis voigti'' (Wichard & Engel, 2006) **Genus ''
Sialis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Latreille, 1802 ***Species Approximately 90 ***Species ''
Sialis abchasica ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis aequalis'' Banks, 1920 ***Species ''
Sialis americana ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' (Rambur, 1842) ***Species '' Sialis annae'' Vshivkova, 1979 ***Species ''
Sialis arvalis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis atra'' Navás, (1928) ***Species ''
Sialis bifida ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Hayashi & Suda, 1997 ***Species ''
Sialis bilineata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Say, 1823 ***Species '' Sialis bilobata'' Whiting, 1991 ***Species ''
Sialis californica ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Banks, 1920 ***Species ''
Sialis chilensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' McLachlan, (1871) ***Species ''
Sialis concava ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Banks, 1897 ***Species '' Sialis contigua'' Flint, 1964 ***Species ''
Sialis cornuta ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis didyma'' Navás, 1916 ***Species ''
Sialis dorochovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis dorsata'' Say, 1823 ***Species ''
Sialis dreisbachi ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Flint, 1964 ***Species '' Sialis elegans'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis flavicollis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Enderlein, 1910 ***Species '' Sialis formosana'' Esben-Petersen, 1913 ***Species '' Sialis frequens'' Okamoto, 1905 ***Species ''
Sialis fuliginosa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' F. Pictet, 1836 ***Species ''
Sialis fumosa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Navás, 1915 ***Species '' Sialis glabella'' Ross, 1937 ***Species ''
Sialis gonzalezi ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species †''
Sialis groehni ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Wichard, 1997 ***Species '' Sialis hamata'' Ross, 1937 ***Species ''
Sialis hasta ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis henanensis'' X.-y. Liu & D. Yang, 2006 ***Species †'' Sialis herrlingi'' Wichard, 2002 ***Species '' Sialis imbecilla'' Say, 1823 ***Species ''
Sialis immarginata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Say, 1823 ***Species ''
Sialis infumata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Newman, 1838 ***Species '' Sialis iola'' Ross, 1937 ***Species ''
Sialis itasca ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species ''
Sialis japonica ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' van der Weele, 1909 ***Species '' Sialis jianfengensis'' D. Yang et al., 2002 ***Species ''
Sialis joppa ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species ''
Sialis klingstedti ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 ***Species '' Sialis kunmingensis'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis levanidovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1980 ***Species ''
Sialis longidens ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Klingstedt, (1932) ***Species '' Sialis lutaria'' (Linnaeus, 1758) ***Species ''
Sialis martynovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1980 ***Species '' Sialis melania'' Nakahara, 1915 ****Subspecies '' Sialis melania kyushuensis'' Hayashi & Suda, 1995 ****Subspecies '' Sialis melania melania'' Nakahara, 1915 ****Subspecies '' Sialis melania tohokuensis'' Hayashi & Suda, 1995 ****Subspecies '' Sialis melania toyamaensis'' Hayashi & Suda, 1995 ***Species '' Sialis mohri'' Ross, 1937 ***Species ''
Sialis morio ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Klingstedt, (1933) ***Species '' Sialis morrisoni'' K. Davis, 1903 ***Species '' Sialis muratensis'' Nel, 1988 ***Species ''
Sialis nevadensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' K. Davis, 1903 ***Species ''
Sialis nigripes ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' E. Pictet, 1865 ***Species '' Sialis nina'' Townsend, 1939 ***Species '' Sialis occidens'' Ross, 1937 ***Species '' Sialis rotunda'' Banks, 1920 ***Species '' Sialis sibirica'' McLachlan, 1872 ***Species ''
Sialis sinensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Banks, (1940) ***Species '' Sialis sordida'' Klingstedt, (1933) ***Species ''
Sialis spangleri ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Flint, 1964 ***Species '' Sialis strausi'' Illies, 1967 ***Species '' Sialis vagans'' Ross, 1937 ***Species ''
Sialis vanderweelei ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' U. Aspöck & H. Aspöck, 1983 ***Species ''
Sialis velata ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Ross, 1937 ***Species '' Sialis versicoloris'' X.-y. Liu & D. Yang, 2006 ***Species ''
Sialis yamatoensis ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Hayashi & Suda, 1995 ***Species ''
Sialis zhiltzovae ''Sialis'' is a genus of alderfly belonging to the order Megaloptera family Sialidae. Description These alderflies are small and mainly brown with a relatively heavy body and forewings reaching a length of 10 to 20 millimeters. Females are us ...
'' Vshivkova, 1985 **Genus '' Stenosialis'' Tillyard, (1919) ***Species '' Stenosialis australiensis'' Tillyard, (1919) ***Species '' Stenosialis hollowayi'' Theischinger, 1983 ''Sialis lutaria'' is the commonest alderfly in the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and ...
(2006)
Alderfly - ''Sialis lutaria''
Retrieved 28 June 2006.
and across much of Europe.


References


External links


BugGuide


{{Authority control Megaloptera Aquatic insects