Shirley M. Tilghman
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Shirley Marie Tilghman, (; née Caldwell; born 17 September 1946) is a Canadian scholar in
molecular biology Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including biomolecular synthesis, modification, mechanisms, and interactions. The study of chemical and physi ...
and an academic administrator. She is now a professor of molecular biology and public policy and president emerita of
Princeton University Princeton University is a private research university in Princeton, New Jersey. Founded in 1746 in Elizabeth as the College of New Jersey, Princeton is the fourth-oldest institution of higher education in the United States and one of the ...
. In 2002, '' Discover'' magazine recognized her as one of the 50 most important women in science. Tilghman was the 19th president of Princeton University; she was the first woman to hold the position and the second female president in the
Ivy League The Ivy League is an American collegiate athletic conference comprising eight private research universities in the Northeastern United States. The term ''Ivy League'' is typically used beyond the sports context to refer to the eight school ...
. Tilghman was also the first biologist to hold the Princeton presidency. She is the fifth foreign-born president of Princeton, and the second academic born in Canada to be elected to the position. A leader in the field of molecular biology, Tilghman was a member of the Princeton faculty for fifteen years before being named president. She has returned to the Princeton faculty as a professor of molecular biology. In that capacity, she has returned to the Lewis-Sigler Institute of Integrative Genomics as a faculty member; while she is not currently engaged in research, Tilghman actively advises undergraduates in their independent research, including the senior thesis for seniors. Tilghman also continues to hold leadership positions in the global scientific community. She was the 2015 president of the
American Society for Cell Biology The American Society for Cell Biology (ASCB) is a professional society that was founded in 1960.Toronto, Ontario Toronto ( ; or ) is the capital city of the Canadian province of Ontario. With a recorded population of 2,794,356 in 2021, it is the most populous city in Canada and the fourth most populous city in North America. The city is the ancho ...
, Canada. As a young child, her father encouraged her interest in math. She graduated from
Kelvin High School Kelvin High School is a public high school in Winnipeg, Manitoba, Canada. The school is located in the neighbourhood of River Heights. Kelvin teaches grades 9 to 12 and is part of the South District of the Winnipeg School Division. History T ...
in
Winnipeg Winnipeg () is the capital and largest city of the province of Manitoba in Canada. It is centred on the confluence of the Red and Assiniboine rivers, near the longitudinal centre of North America. , Winnipeg had a city population of 749, ...
,
Manitoba , image_map = Manitoba in Canada 2.svg , map_alt = Map showing Manitoba's location in the centre of Southern Canada , Label_map = yes , coordinates = , capital = Winn ...
and received her honours B.Sc. in chemistry from Queen's University in
Kingston, Ontario Kingston is a city in Ontario, Canada. It is located on the north-eastern end of Lake Ontario, at the beginning of the St. Lawrence River and at the mouth of the Cataraqui River (south end of the Rideau Canal). The city is midway between To ...
, in 1968. She was a secondary school teacher in
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierr ...
, West Africa, in the Canadian University Services Overseas (CUSO) Program. Tilghman earned her Ph.D. in biochemistry from
Temple University Temple University (Temple or TU) is a public state-related research university in Philadelphia, Pennsylvania. It was founded in 1884 by the Baptist minister Russell Conwell and his congregation Grace Baptist Church of Philadelphia then calle ...
in
Philadelphia, Pennsylvania Philadelphia, often called Philly, is the largest city in the Commonwealth of Pennsylvania, the sixth-largest city in the U.S., the second-largest city in both the Northeast megalopolis and Mid-Atlantic regions after New York City. Sinc ...
under Richard W. Hanson. Tilghman was Hanson's first graduate student. Her PhD Dissertation was entitled ''"The Hormonal Regulation of Phosphoenolpyruvate Carboxykinase."''


Personal life

She married Joseph Tilghman in 1970. This marriage ended in 1983, leaving Shirley Tilghman with custody of their young daughter (Rebecca) and infant son (Alex). She attributes her successful balancing of a scientific career and caring for her family to organization and focus. Her goal was to not feel guilty while at work or at home, instead focusing on the task at hand.


Research

Tilghman's work in
molecular genetics Molecular genetics is a sub-field of biology that addresses how differences in the structures or expression of DNA molecules manifests as variation among organisms. Molecular genetics often applies an "investigative approach" to determine the ...
focused on the regulation of genes during development, particularly in the field of genomic imprinting. During postdoctoral studies at the
National Institutes of Health The National Institutes of Health, commonly referred to as NIH (with each letter pronounced individually), is the primary agency of the United States government responsible for biomedical and public health research. It was founded in the late ...
, Tilghman made a number of discoveries while a member of the team which cloned the first mammalian gene. She went on to demonstrate that the
globin The globins are a superfamily of heme-containing globular proteins, involved in binding and/or transporting oxygen. These proteins all incorporate the globin fold, a series of eight alpha helical segments. Two prominent members include myogl ...
gene was spliced, a finding that helped confirm some of the revolutionary theories then emerging about gene behavior. As an independent investigator at the
Fox Chase Cancer Center Fox Chase Cancer Center is a National Cancer Institute-designated Comprehensive Cancer Center research facility and hospital located in the Fox Chase section of Philadelphia, Pennsylvania, United States. The main facilities of the center are loca ...
in Philadelphia from 1980 to 1986 and adjunct associate professor of Human Genetics at the
University of Pennsylvania The University of Pennsylvania (also known as Penn or UPenn) is a private research university in Philadelphia. It is the fourth-oldest institution of higher education in the United States and is ranked among the highest-regarded universitie ...
Tilghman continued to make scientific breakthroughs. Tilghman went to Princeton University in 1986 as the Howard A. Prior Professor of the Life Sciences. Two years later, she also joined the Howard Hughes Medical Institute as an investigator. She was a leader in the use of mice to understand the behavior of genes by researching the effect of gene insertion in embryonic cells. In 1998, she took on additional responsibilities as the founding director of Princeton's multi-disciplinary Lewis-Sigler Institute for Integrative Genomics, while continuing to study how male and female genomes are packaged and the consequences of the differences for regulating embryo growth. Tilghman's extensive series of published research papers are catalogued on the
PubMed PubMed is a free search engine accessing primarily the MEDLINE database of references and abstracts on life sciences and biomedical topics. The United States National Library of Medicine (NLM) at the National Institutes of Health maintain t ...
government website of the
United States National Library of Medicine The United States National Library of Medicine (NLM), operated by the United States federal government, is the world's largest medical library. Located in Bethesda, Maryland, the NLM is an institute within the National Institutes of Health. Its ...
, the NLM division of the
National Institutes of Health The National Institutes of Health, commonly referred to as NIH (with each letter pronounced individually), is the primary agency of the United States government responsible for biomedical and public health research. It was founded in the late ...
.


President of Princeton University

Tilghman succeeded Harold Tafler Shapiro and became the 19th president of Princeton University in 2001. She was elected Princeton's first woman president on May 5, 2001, and assumed office on June 15, 2001. Under her administration, the university built a sixth residential college, named in honor of alumna
Meg Whitman Margaret Cushing Whitman (born August 4, 1956) is the US ambassador to Kenya, an American business executive and former gubernatorial candidate for California. She is a board member of Procter & Gamble and General Motors. Whitman was previously ...
, to accommodate an 11 percent expansion of the undergraduate student body (an increase of some 500 students), as recommended by a special committee of the Board of Trustees chaired by Paul M. Wythes. In 2012, Tilghman announced that she would step down from her presidency in June 2013. She was succeeded by the university's then-provost, Christopher L. Eisgruber. For Tilghman, Princeton has two essential missions. "One is to ensure that our doors are open as wide as possible to every talented student in the world who is capable of doing the hard work we ask of them. And that means maintaining our commitment to financial aid, which is the tool – the critical tool – to get those students to Princeton. And the second thing is that we must address the most critical issues, and push back the frontiers of knowledge, and not just in science and technology, but in social policy, and in public policy, and in understanding the nature of the human condition." The establishment of
Whitman College Whitman College is a private liberal arts college in Walla Walla, Washington. The school offers 53 majors and 33 minors in the liberal arts and sciences, and it has a student-to-faculty ratio of 9:1. Whitman was the first college in the Pacific ...
, together with the reconstruction of Butler College, accompanied a significant reconfiguration of Princeton's residential college system, which now incorporates upperclassmen as well as freshmen and sophomores, providing new residential options and increasing opportunities for social interaction across the student body. In addition, an effort has been made to strengthen the relationship between the university and Princeton's independent eating clubs, where most upperclassmen take their meals, with the goal of enhancing the undergraduate experience of all students. In 2009, she appointed a committee chaired by Nannerl O. Keohane to review undergraduate women's leadership at Princeton.


Academics

Tilghman has presided over a number of academic initiatives at Princeton, including the creation of a Center for African American Studies, the Lewis Center for the Arts (named after alumnus Peter B. Lewis), the Princeton Neuroscience Institute and the Andlinger Center for Energy and the Environment (after alumnus Gerhard R. Andlinger). Along with the renewal of the Department of Chemistry, these steps have both capitalized on Princeton's existing strengths and broken new ground, ensuring that the university will, in Tilghman's words, continue ''"to make the world a better place through the power of the mind and the imagination."''


Diversity

More broadly, Tilghman's presidency has placed an emphasis on increasing the diversity of Princeton's faculty and students; widening access to the university through improvements to its generous financial aid program and the elimination of admission through "early decision"; fostering a multidisciplinary approach to teaching and research; and strengthening the university's international perspective through a wide range of initiatives – from the Global Scholars Program, which brings international scholars to campus on a recurring basis, to the Bridge Year Program, which gives incoming freshmen an opportunity to defer their studies for a year in order to devote themselves to public service overseas.


Funding higher education


Student loans

As her presidency started the university accomplished the long-hoped-for goal of eliminating the need for student loans; Princeton became the first American university to replace student loans with grants from its endowment. In principle, students earning a Princeton degree could graduate debt free.


Tuition

The size of the endowment and the success of these programs prompted some to question whether Tilghman would implement a policy of eliminating tuition altogether. In her ''Wall Street Journal'' article on this matter, she indicated that Princeton would continue to charge tuition, and that she felt that charging tuition was a morally and economically correct policy to maintain.


Fundraising

During her tenure the percentage of students receiving some form of financial aid increased and the size of the average award also increased. These policies were partially facilitated by the growing size of the university's financial endowment, whose income is used to finance the university's mission alongside tuition, and the annual funding of the operating budget through alumni donations from Princeton's Annual Giving campaign.


Controversies


Pro-women hirings

Although President Tilghman has been accused of favoring women in her hiring practices, in fact, most of her appointees have been men. The women she has hired to senior positions include
Amy Gutmann Amy Gutmann (born November 19, 1949) is an American academic and diplomat who is the United States Ambassador to Germany. She was the eighth president of the University of Pennsylvania. In November 2016, the school announced that her contract ...
as provost, the second-most-powerful administrative position in the university,
Anne-Marie Slaughter Anne-Marie Slaughter (born September 27, 1958) is an American international lawyer, foreign policy analyst, political scientist and public commentator. From 2002 to 2009, she was the Dean of Princeton University's School of Public and Interna ...
as Dean of the
Woodrow Wilson School of Public and International Affairs The Princeton School of Public and International Affairs (formerly the Woodrow Wilson School of Public and International Affairs) is a professional public policy school at Princeton University. The school provides an array of comprehensive course ...
as well as her successor Christina Paxson,
Maria Klawe Maria Margaret Klawe ( ; born 1951) is a computer scientist and the fifth president of Harvey Mudd College (since July 1, 2006). Born in Toronto in 1951, she became a naturalized U.S. citizen in 2009. She was previously Dean of the School of En ...
as Dean of the School of Engineering and Applied Science, and Janet Lavin Rapelye as the Dean of Admission. Gutman would go on to lead the
University of Pennsylvania The University of Pennsylvania (also known as Penn or UPenn) is a private research university in Philadelphia. It is the fourth-oldest institution of higher education in the United States and is ranked among the highest-regarded universitie ...
as their president in early 2004; Klawe was chosen president of Harvey Mudd College in 2006. Slaughter took a leave from the university to serve as Director of Policy and Planning at the U.S. State Department, reporting to the Secretary of State at the time,
Hillary Clinton Hillary Diane Rodham Clinton ( Rodham; born October 26, 1947) is an American politician, diplomat, and former lawyer who served as the 67th United States Secretary of State for President Barack Obama from 2009 to 2013, as a United States sen ...
. Paxson became president of Brown University in 2012. Tilghman has appointed prominent men to leadership positions at Princeton, such as Charles Kalmbach as the senior vice president for administration, the highest non-academic administrative post, David P. Dobkin as dean of the faculty, Gutmann's replacement as provost and Tilghman's successor Christopher L. Eisgruber, and Klawe's replacement
Vincent Poor Vincent ( la, Vincentius) is a male given name derived from the Roman name Vincentius, which is derived from the Latin word (''to conquer''). People with the given name Artists *Vincent Apap (1909–2003), Maltese sculptor *Vincent van Gogh ...
. She initiated a review of undergraduate women's leadership at Princeton, chaired by Nannerl O. Keohane; the review found that the early prominence for women in leadership positions that accompanied to introduction of women students to the campus had recently not been as frequently repeated.


Ivy League seven-week athletic moratorium

Tilghman signed on to the
Ivy League The Ivy League is an American collegiate athletic conference comprising eight private research universities in the Northeastern United States. The term ''Ivy League'' is typically used beyond the sports context to refer to the eight school ...
-wide seven-week athletic moratorium, since modified, in which intercollegiate athletes were enjoined from taking part in supervised practices and other obligatory athletic activities for seven weeks during the academic year in order to encourage them to participate in other activities. Supporters of the proposal pointed to studies by former Princeton president William G. Bowen, whose controversial book ''The Game of Life'' purported to describe widespread academic "underperformance" of college athletes. Detractors claimed that the book was flawed, and the moratorium represented an encroachment on students' freedom to use their time as they saw fit.


University donor complaint

During her presidency, Tilghman was embroiled in a court case pitting her against the family of a major donor to the university, the Robertsons. The children of the original donors, who were themselves Princeton alums, alleged that the university failed to apply the funds donated by their parents to the intended purpose, and asked for the funds to be restored to the family for use elsewhere. The donated funds had become joined with the university's general endowment, resulting in efficiencies and benefits in fund management and performance, which were not the subject of the Robertsons' complaints. The case was settled in 2008 with a payment to the family of $40 million in legal fees and another $50 million, plus interest, to a new foundation to support education for government service. Princeton retained the remainder of the money for the Wilson school.


Corporate board matters

In August 2012, Tilghman was
subpoena A subpoena (; also subpœna, supenna or subpena) or witness summons is a writ issued by a government agency, most often a court, to compel testimony by a witness or production of evidence under a penalty for failure. There are two common types of ...
ed in her dual capacities as Princeton's president and as a member of Google's board, as part of a suit to block a board approved 2-for-1 Google stock split that the complaining party claimed would represent "an unfair effort to diminish its voting power while reserving voting rights for the company’s founders
Larry Page Lawrence Edward Page (born March 26, 1973) is an American business magnate, computer scientist and internet entrepreneur. He is best known for co-founding Google with Sergey Brin. Page was the chief executive officer of Google from 1997 unti ...
and
Sergey Brin Sergey Mikhailovich Brin (russian: link=no, Сергей Михайлович Брин; born August 21, 1973) is an American business magnate, computer scientist, and internet entrepreneur, who co-founded Google with Larry Page. Brin was th ...
". All members of the Google board received subpoenas. In the case of Tilghman, "records of donations, contribution pledges or promises made by Tilghman or the university to any charities, organizations, foundations or educational institutions that have any affiliation with Page, Brin, Schmidt or Google" were demanded.


Precedents

While Tilghman disquieted some alumni by championing affirmative action policies, establishing a single admission process, and broadening the range of residential and dining options available to students, she also found strong support for these actions and their underpinning vision. Tilghman presided over a major effort to advance the growing community of Princeton Alumnae, culminating in a campus conference entitled "She Roars". In her final year, Tilghman led the first major university celebration for "alternative genders", resulting in an immensely successful on-campus LGBT alumni gathering; this was the first of its kind on any campus in the United States and set a precedent for the advancement of the LGBT community nationwide.


Internal roles

As president of Princeton University, Tilghman was also an ex-officio trustee of the Princeton Board of Trustees, and chairman of Princeton Honorary Degrees Committee. Tilghman orchestrated an orderly transition from her other Princeton roles after being named president.
David Botstein David Botstein (born September 8, 1942) is an American biologist serving as the chief scientific officer of Calico. He served as the director of the Lewis-Sigler Institute for Integrative Genomics at Princeton University from 2003 to 2013, where ...
succeeded Tilghman as the Director of the Lewis-Sigler Institute of Integrative Genomics in 2003. Ned S Wingreen succeeded her as the Howard A Prior Professor of Life Sciences. She eventually closed her lab to dedicate her time fully to the presidency; in doing so, she assured that all students who had begun with her as adviser were able to successfully complete their degrees and associated research before the lab closed. An award-winning teacher, Tilghman continued to conduct classes even while serving as president. She became a "Princeton Parent" when her daughter matriculated at Princeton as an undergraduate during her tenure as president and was accorded honorary alumna status to a record number of Princeton classes.


External roles

While serving as president of Princeton, Tilghman accepted membership on the board of directors of
Google Google LLC () is an American Multinational corporation, multinational technology company focusing on Search Engine, search engine technology, online advertising, cloud computing, software, computer software, quantum computing, e-commerce, ar ...
, and served in that capacity from October 2005 to February 2018. As compensation for joining the board, she received 6,000 shares of stock that by 2005 were worth in excess of her Princeton compensation package that by 2003 had reached $533,057. Beginning in 2001, she served for a time on the Queen University's Chemistry Innovation Council in order to help the development of the Chemistry program at Queen's, which is based in
Kingston, Ontario Kingston is a city in Ontario, Canada. It is located on the north-eastern end of Lake Ontario, at the beginning of the St. Lawrence River and at the mouth of the Cataraqui River (south end of the Rideau Canal). The city is midway between To ...
, Canada. Tilghman served on the board of trustees at
Cold Spring Harbor Laboratory Cold Spring Harbor Laboratory (CSHL) is a private, non-profit institution with research programs focusing on cancer, neuroscience, plant biology, genomics, and quantitative biology. It is one of 68 institutions supported by the Cancer Centers ...
(CSHL), Long Island, New York, during the early years of her presidency. In 2006, Tilghman was one of three sitting university presidents who served on the Duke University President's Council that investigated the university's wide-ranging actions after the lacrosse players scandal wherein members of the Duke lacrosse team were charged with various types of inappropriate and allegedly illegal off campus behavior.


Successor

On September 21, 2012, Shirley informed the Princeton Board of Trustees that she planned to step down as the 19th president of Princeton University at the end of the 2012 academic year. On April 21, 2013, it was announced that Christopher L. Eisgruber would succeed Tilghman as Princeton's president, effective July 1. Notable Princeton alumnus Peter Lewis 1955 said at the time, "Ideally, she'd be remembered for grooming a terrific successor."


Awards and honors


Memberships

Tilghman has been elected to the following organizations: * Member of the
American Philosophical Society The American Philosophical Society (APS), founded in 1743 in Philadelphia, is a scholarly organization that promotes knowledge in the sciences and humanities through research, professional meetings, publications, library resources, and communit ...
(2000), *Foreign Associate of the National Academy of Sciences, with Primary Field ''Cellular and Developmental Biology'' and Secondary Field ''Genetics'' *Member, the Institute of Medicine *Elected a
Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the judges of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural knowledge, including mathemat ...
(1995) * Founding member of the
International Mammalian Genome Society The International Mammalian Genome Society (IMGS) is a professional scientific organization that promotes and coordinates the genetic and genomic study of mammals. It has a scientific journal, ''Mammalian Genome'', and organizes an annual interna ...
. *Honorary Member, Institute of Electrical and Electronics Engineers (IEEE) (2014) She serves as a trustee of the Carnegie Endowment for International Peace and the
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
’s
King Abdullah University of Science and Technology King Abdullah University of Science and Technology (KAUST; ar, جامعة الملك عبد الله للعلوم و التقنية ') is a private research university located in Thuwal, Saudi Arabia. Founded in 2009, the university provid ...
. From 1993 through 2000, Tilghman chaired Princeton's Council on Science and Technology, which encourages teaching science and technology to students outside the sciences. In 1996, she received Princeton's President's Award for Distinguished Teaching. Nineteen Princeton graduating classes, from 1941 to 2005, have made President Tilghman an honorary member.


Awards

Tilghman has earned the following awards during her career: *
American Society for Cell Biology The American Society for Cell Biology (ASCB) is a professional society that was founded in 1960. * L'Oréal-UNESCO Awards for Women in Science (2002) * Lifetime Achievement Award from the Society for Developmental Biology (2003) * The Memorial Sloan-Kettering Medal for Outstanding Contributions to Biomedical Research (2005) * Genetics Society of America Medal (2007) * American Dream Award to Successful Immigrants to the U.S.A. (2007) * Henry G. Friesen International Prize in Health Research (2010) * Officer of the
Order of Canada The Order of Canada (french: Ordre du Canada; abbreviated as OC) is a Canadian state order and the second-highest honour for merit in the system of orders, decorations, and medals of Canada, after the Order of Merit. To coincide with the cen ...
(2014) * Benjamin Franklin Creativity Laureate in Science and Public Service, the Creativity Foundation (2014) * Person of the Year, Science Careers from the Journal of Science (2014) *
UCSF The University of California, San Francisco (UCSF) is a public land-grant research university in San Francisco, California. It is part of the University of California system and is dedicated entirely to health science and life science. It con ...
(2017). *
Patrusky Lecture The Patrusky lecture series is held by the Council for the Advancement of Science Writing (CASW) in honor of Ben Patrusky, who retired from CASW in 2013 after 25 years as the executive director of CASW and 30 years as the director of the New Horizon ...
r (2018) * Helen Dean King Award (2018)


Honorary degrees

The 19th president of Princeton University has received more than 19 honorary academic degrees: * Honorary Doctor of Science, Bard College (2002) * Honorary degree, Dickinson College (2002) * Honorary Doctor of Laws, Simon Fraser University (2002) * Honorary degree, University of British Columbia (2002) * Honorary Doctor of Science, Yale University (2002) * Honorary Doctor of Laws, Rider University (2002) * Honorary degree, Drew University (2004) * Honorary Doctor of Laws, Harvard University (2004) * Honorary degree, Watson School of Biological Sciences, Cold Spring Harbor Laboratories (2004) * Honorary Doctor of Science, Columbia University (2005) * Honorary Doctor of Science, New York University (2005) * Honorary Doctorate, Rockefeller University (2006) * Honorary Doctor of Science, Rutgers University, The State University of New Jersey (2006) * Honorary degree, Memorial University of Newfoundland (2007) * Honorary Doctorate, Ryerson University (2007) * Honorary Doctor of Science, Washington University (2007) * Honorary Doctor of Human Letters, Amherst College (2008) * Honorary degree, Rensselaer (RPI) (2008) * Honorary Doctor of Science, UMBC (2009) * Honorary Doctor of Laws, Princeton University (2013)


Roles after Princeton presidency

Tilghman continues as a member of the Princeton faculty in the Department of Molecular Biology and is a faculty member of the Lewis-Sigler Institute for Integrative Genomics. Upon leaving the Princeton presidency, Tilghman retained her seat on the Google board. At the time, Google's Eric Schmidt supported this retention by emphasizing the benefits Google had received from Tilghman generally in her service on the Board. Tilghman continues to serve as a Trustee of Amherst College. She is a member of the board of the Brookhaven Science Associates, the organization that manages
Brookhaven National Laboratory Brookhaven National Laboratory (BNL) is a United States Department of Energy national laboratory located in Upton, Long Island, and was formally established in 1947 at the site of Camp Upton, a former U.S. Army base and Japanese internment c ...
on Long Island, New York. She is a Trustee of the
Institute for Advanced Study The Institute for Advanced Study (IAS), located in Princeton, New Jersey, in the United States, is an independent center for theoretical research and intellectual inquiry. It has served as the academic home of internationally preeminent schola ...
. Tilghman serves on the board of the Broad Institute, founded to encourage a unique model of collaborative, inter-institutional research, initially through joint efforts between Harvard and MIT. She is a Trustee of LEDA, the Leadership Enterprise for a Diverse America. In 2013, Tilghman was elected to serve as the 2015 president of the ASCB


Quotes

"What made it truly thrilling was that the genes were organized in a way that was totally unexpected. So nature took us by surprise." "There are 25 years of good social science that demonstrate the many cultural practices that act collectively to discourage women from entering and continuing careers in science and engineering. The research is overwhelming, and it is there for anybody to see. On the other hand, the data that would suggest there are innate differences in the abilities of men and women to succeed in the natural sciences are nonexistent." On how she hoped to spend her time during her sabbatical before returning to the faculty in 2014, Tilghman said, "I’m going to be an attentive grandmother." (In ''
The Daily Princetonian ''The Daily Princetonian'', originally known as ''The Princetonian'' and nicknamed the Prince, is the independent daily student newspaper of Princeton University. Founded on June 14, 1876 as ''The'' ''Princetonian'', it changed its name to ''T ...
'' article ''"Princeton’s biggest fan, Princeton’s biggest critic"'' by Luc Cohen, May 30, 2013.) On explaining the thinking that led her to found a backup day-care option for Princeton employees introduced in 2007, an idea that reflected her own career management while raising two children, Tilghman said, "For years, I had said that when I retired, I would start a company that hired retired people to sit for working mothers..."


Publications

Tilghman's publications as a research scientist are referenced in the Research section. Her other major publications are as follows: * Tilghman, Shirley, et al. (1994) The Funding of Young Investigators in the Biological and Biomedical Sciences. * Tilghman, Shirley and National Research Council Committee on Dimensions, Causes, and Implications of Recent Trends in the Careers of Life Scientists (1998) Trends in the Careers of Life Scientists
Molecular Biology of the Cell Vol. 9, 3007–3015
* Tilghman, Shirley (1999) The Sins of the Fathers and Mothers: Genomic Imprinting in Mammalian Development
Cell Volume 96, Issue 2
* Tilghman, Shirley, et al. (2014) Rescuing US Biomedical Research from its Systemic Flaws; Proceedings of the National Academy of Sciences of America (PNAS), vol. 111, no. 16 Tilghman wrote the "President's Page" in each edition of the Princeton Alumni Weekly (the PAW) during her term as president. Electronic archives of past PAW issues are available at the magazine's website.


Speeches

Tilghman delivered eleven Commencement addresses at Princeton University during her tenure as president. The text of each Princeton Commencement Address is available at the Tilghman e-Archive on the website of the office of the president at Princeton University. She has spoken at a number of other commencement and graduation exercises, often when awarded an honorary degree. Tilghman has often been asked to deliver commemorative speeches, sometimes known as "Remarks", give testimony, and participate in panels at other universities and notable venues, including: * The Dehejia Lecture at the Sidwell Friends School, as the Dehejia Fellow: "Science: The Last Frontier." (2006) * The Ullyot Public Affairs Lecture at the Chemical Heritage Foundation: "Strategy or Happenstance:Science Policy in the U.S.A."(2007) * The Baldwin Lecture at Princeton University: "The Meaning of Race in the Post-Genome Era" (2010) * The inauguration of Anthony Monaco as president of
Tufts University Tufts University is a private research university on the border of Medford and Somerville, Massachusetts. It was founded in 1852 as Tufts College by Christian universalists who sought to provide a nonsectarian institution of higher learning. ...
(2011) * Testimony before the U.S. Congress on Immigration and Visas She has delivered the following additional addresses. * Tilghman, Shirley (2005) Recruiting, Retaining and Advancing Women Scientists in Academia
Address delivered March 24 at Columbia University


References


External links


A Tilghman timeline


* ttps://www.princeton.edu/pr/smt/bio.html Tilghman's biography from Princeton
Office of the President of Princeton University

Shirley Tilghman
Video produced by '' Makers: Women Who Make America''
Shirley Tilghman iBiology Seminar: The Malthusian Dilemma in Biomedical Research
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