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Shearwaters are medium-sized long-winged
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s in the
petrel Petrels are tube-nosed seabirds in the bird order Procellariiformes. Description The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the albatross f ...
family
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which als ...
. They have a global marine distribution, but are most common in temperate and cold waters, and are
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
outside the breeding season.


Description

These tubenose birds fly with stiff wings and use a "shearing" flight technique (flying very close to the water and seemingly cutting or "shearing" the tips of waves) to move across wave fronts with the minimum of active flight. This technique gives the group its English name. Some small species, like the Manx shearwater are
cruciform Cruciform is a term for physical manifestations resembling a common cross or Christian cross. The label can be extended to architectural shapes, biology, art, and design. Cruciform architectural plan Christian churches are commonly describe ...
in flight, with their long wings held directly out from their bodies.


Behaviour


Movements

Many shearwaters are long-distance migrants, perhaps most spectacularly
sooty shearwater The sooty shearwater (''Ardenna grisea'') is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name , and as muttonbird, like its relatives the wedge-tailed shearwater (''A. pacificus ...
s, which cover distances in excess of from their breeding colony on the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouze ...
(52°S 60°W) to as far as 70° north latitude in the North Atlantic Ocean off northern Norway. One study found Sooty shearwaters migrating nearly a year, which would give them the longest animal migration ever recorded electronically. Short-tailed shearwaters perform an even longer "figure of eight" loop migration in the Pacific Ocean from
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
to as far north as the Arctic Ocean off northwest Alaska. They are long-lived. A Manx shearwater breeding on Copeland Island, Northern Ireland, was (as of 2003/2004) the oldest known wild bird in the world: ringed as an adult (when at least 5 years old) in July 1953, it was retrapped in July 2003, at least 55 years old. Manx shearwaters migrate over to South America in winter, using waters off southern Brazil and Argentina, so this bird had covered a minimum of on migration alone. Following the tracks of the migratory
Yelkouan shearwater upright=0.8, Egg of the yelkouan shearwater The yelkouan shearwater, Levantine shearwater or Mediterranean shearwater (''Puffinus yelkouan'') is a medium-sized shearwater in the seabird family Procellariidae. It was formerly treated as a subsp ...
has revealed that this species never flies overland, even if it means flying an extra 1'000 km. For instance, during their seasonal migration towards the Black Sea they would circumvent the entire Peloponnese instead of crossing over the 6 km isthmus of Corinth.


Breeding

Shearwaters come to islands and coastal cliffs only to breed. They are nocturnal at the colonial breeding sites, preferring moonless nights to minimize predation. They nest in burrows and often give eerie contact calls on their night-time visits. They lay a single white egg. The chicks of some species, notably short-tailed and sooty shearwaters, are subject to harvesting from their nest burrows for food, a practice known as muttonbirding, in Australia and New Zealand.


Feeding

They feed on fish, squid, and similar oceanic food. Some will follow fishing boats to take scraps, commonly the sooty shearwater; these species also commonly follow
whale Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and ...
s to feed on fish disturbed by them. Their primary feeding technique is diving, with some species diving to depths of .


Taxonomy

There are about 30
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
: a few larger ones in the genera ''
Calonectris ''Calonectris'' is a genus of seabirds. The genus name comes from Ancient Greek ''kalos'', "good" and ''nectris'', "swimmer". The genus comprises four large shearwaters. There are two other shearwater genera. ''Puffinus'', which comprises abou ...
'' and ''
Ardenna ''Ardenna'' is a genus of seabirds in the family Procellariidae. These medium-sized shearwater species were formerly included in the genus ''Puffinus''. Taxonomy A phylogenetic analysis using mitochondrial DNA published in 2004 found that ''Puff ...
'' and many smaller ones in ''
Puffinus ''Puffinus'' is a genus of seabirds in the order Procellariiformes that contains about 20 small to medium-sized shearwaters. Two other shearwater genera are named: '' Calonectris'', which comprises three or four large shearwaters, and '' Ardenna' ...
''. Recent genomic studies show that Shearwaters form a clade with ''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The specta ...
'', ''
Bulweria ''Bulweria'' is a genus of seabirds in the family Procellariidae named after English naturalist James Bulwer. The genus has two extant species, Bulwer's petrel and Jouanin's petrel . A third species, the Olson's petrel, became extinct in the ...
'' and ''
Pseudobulweria __NOTOC__ ''Pseudobulweria'' is a genus of seabirds in the family Procellariidae. They have long been retained with the gadfly petrel genus ''Pterodroma'' despite morphological differences. Mitochondrial DNA cytochrome ''b'' sequence analysis ...
''. This arrangement contrasts with earlier conceptions based on mitochondrial DNA sequencing.


List of species

The group contains 3 genera with 32 species. * ''
Puffinus ''Puffinus'' is a genus of seabirds in the order Procellariiformes that contains about 20 small to medium-sized shearwaters. Two other shearwater genera are named: '' Calonectris'', which comprises three or four large shearwaters, and '' Ardenna' ...
'' **
Christmas shearwater The Christmas shearwater or ''aoū'' (''Puffinus nativitatis'') is a medium-sized shearwater of the tropical Central Pacific. It is a poorly known species due to its remote nesting habits, and it has not been extensively studied at sea either. ...
, ''Puffinus nativatis'' ** Manx shearwater, ''Puffinus puffinus'' **
Yelkouan shearwater upright=0.8, Egg of the yelkouan shearwater The yelkouan shearwater, Levantine shearwater or Mediterranean shearwater (''Puffinus yelkouan'') is a medium-sized shearwater in the seabird family Procellariidae. It was formerly treated as a subsp ...
, ''Puffinus yelkouan'' **
Balearic shearwater The Balearic shearwater (''Puffinus mauretanicus'') is a medium-sized shearwater in the seabird family ''Procellariidae''. ''Puffinus'' is a New Latin loanword based on the English "puffin" and its variants, that referred to the cured carcass of ...
, ''Puffinus mauretanicus'' ** Bryan's shearwater, ''Puffinus bryani'' – first described in 2011 **
Black-vented shearwater The black-vented shearwater (''Puffinus opisthomelas'') is a species of seabird. The bird is 30–38 cm in length with a 76–89 cm wingspan. Formerly considered a subspecies of the Manx shearwater, its actual taxonomic relationship ...
, ''Puffinus opisthomelas'' ** Townsend's shearwater, ''Puffinus auriculatus'' ** Newell's shearwater, ''Puffinus newelli'' (split from Townsend's shearwater) ** Rapa shearwater, ''Puffinus myrtae'' (split from Newell's shearwater) ** Fluttering shearwater, ''Puffinus gavia'' ** Hutton's shearwater, ''Puffinus huttoni'' ** Audubon's shearwater, ''Puffinus lherminieri'' ** Persian shearwater, ''Puffinus persicus'' (split from Audubon's shearwater) **
Tropical shearwater The tropical shearwater (''Puffinus bailloni'') is a seabird in the family Procellariidae formerly considered conspecific with Audubon's shearwater (''Puffinus lherminieri''). Subspecies There are five listed subspecies of the tropical shearwa ...
, ''Puffinus bailloni'' (split from Audubon's shearwater) ** Galápagos shearwater, ''Puffinus subalaris'' (split from Audubon's shearwater) ** Bannerman's shearwater, ''Puffinus bannermani'' **
Heinroth's shearwater Heinroth's shearwater (''Puffinus heinrothi'') is a poorly known seabird in the family Procellariidae. Probably a close relative of the little shearwater or Audubon's shearwater (with which it is sometimes considered conspecific), it is distingu ...
, ''Puffinus heinrothi'' **
Little shearwater The little shearwater (''Puffinus assimilis'') is a small shearwater in the petrel family Procellariidae. Despite the generic name, it is unrelated to the puffins, which are auks, the only similarity being that they are both burrow-nesting s ...
, ''Puffinus assimilis'' **
Subantarctic shearwater The subantarctic shearwater (''Puffinus elegans'') is a small bird species which breeds in Tristan da Cunha, islands of the southern Indian Ocean and New Zealand Subantarctic Islands. Taxonomy The subantarctic shearwater is sometimes considered ...
, ''Puffinus elegans'' (split from little shearwater) ** Barolo shearwater or Macronesian shearwater, ''Puffinus baroli'' ** Boyd's shearwater, ''Puffinus boydi'' (split from Barolo shearwater) * ''
Calonectris ''Calonectris'' is a genus of seabirds. The genus name comes from Ancient Greek ''kalos'', "good" and ''nectris'', "swimmer". The genus comprises four large shearwaters. There are two other shearwater genera. ''Puffinus'', which comprises abou ...
'' **
Streaked shearwater The streaked shearwater (''Calonectris leucomelas'') is a species of seabird. The adult bird averages in length, with a wingspan. Description The streaked shearwater feeds mainly on fish and squid. It follows fishing boats, attracted to anch ...
, ''Calonectris leucomelas'' **
Scopoli's shearwater Scopoli's shearwater (''Calonectris diomedea'') is a seabird in the petrel family Procellariidae. It breeds on rocky islands and on steep coasts in the Mediterranean but outside the breeding season it forages in the Atlantic. It is brownish grey ...
, ''Calonectris diomedea'' (split from Cory's shearwater) **
Cory's shearwater Cory's shearwater (''Calonectris borealis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially of rocky islands in the eastern Atlantic. Outside the breeding season it ranges widely in the Atlantic. It was formerl ...
, ''Calonectris diomedea'' **
Cape Verde shearwater The Cape Verde shearwater (''Calonectris edwardsii''), or cagarra locally, is a medium-large shearwater, a seabird in the petrel family Procellariidae. It is Endemism in birds, endemic to the Cape Verde archipelago of Macaronesia in the Atlantic ...
, ''Calonectris edwardsii'' * ''
Ardenna ''Ardenna'' is a genus of seabirds in the family Procellariidae. These medium-sized shearwater species were formerly included in the genus ''Puffinus''. Taxonomy A phylogenetic analysis using mitochondrial DNA published in 2004 found that ''Puff ...
'' **
Wedge-tailed shearwater The wedge-tailed shearwater (''Ardenna pacifica'') is a medium-large shearwater in the seabird family Procellariidae. It is one of the shearwater species that is sometimes referred to as a muttonbird, like the sooty shearwater of New Zealand and ...
, ''Ardenna pacifica'' ** Buller's shearwater, ''Ardenna bulleri'' **
Sooty shearwater The sooty shearwater (''Ardenna grisea'') is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name , and as muttonbird, like its relatives the wedge-tailed shearwater (''A. pacificus ...
, ''Ardenna grisea'' ** Short-tailed shearwater or mutton bird, ''Ardenna tenuirostris'' ** Pink-footed shearwater, ''Ardenna creatopus'' **
Flesh-footed shearwater The flesh-footed shearwater (''Ardenna carneipes''; formerly ''Puffinus carneipes'') is a medium-sized shearwater. Its plumage is black. It has pale pinkish feet, and a pale bill with a distinct black tip. Together with the equally light-billed p ...
, ''Ardenna carneipes'' **
Great shearwater The great shearwater (''Ardenna gravis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic. Taxonomy The great s ...
, ''Ardenna gravis'' There are two extinct species that have been described from fossils. * †
Lava shearwater The lava shearwater (''Puffinus olsoni''), or Olson's shearwater, was a species of shearwater that bred on Lanzarote and Fuerteventura in the Canary Islands. It is known from fossil remains, and was only described in 1990. It was intermediate ...
or Olson's shearwater, ''Puffinus olsoni'' * † Dune shearwater or Hole's shearwater, ''Puffinus holeae''


Phylogeny

Phylogeny of the shearwaters based on a study by Joan Ferrer Obiol and collaborators published in 2022. Only 14 of the 21 recognised species in the genus ''
Puffinus ''Puffinus'' is a genus of seabirds in the order Procellariiformes that contains about 20 small to medium-sized shearwaters. Two other shearwater genera are named: '' Calonectris'', which comprises three or four large shearwaters, and '' Ardenna' ...
'' were included.


References

{{reflist, refs= {{cite journal , last1=Bretagnolle , first1=Vincent , last2=Attié , first2=Carole , last3=Pasquet , first3=Eric , year=1998 , title=Cytochrome-''B'' evidence for validity and phylogenetic relationships of ''Pseudobulweria'' and ''Bulweria'' (Procellariidae) , journal=The Auk , doi=10.2307/4089123 , volume=115 , issue=1 , pages=188–195 , url=http://sora.unm.edu/sites/default/files/journals/auk/v115n01/p0188-p0195.pdf , jstor=4089123 {{cite journal , last1=Nunn , first1=Gary B. , last2=Stanley , first2=Scott E. , year=1998 , title= Body Size Effects and Rates of Cytochrome ''b'' Evolution in Tube-Nosed Seabirds , journal=Molecular Biology and Evolution , doi=10.1093/oxfordjournals.molbev.a025864 , pmid=9787440 , volume=15 , issue=10 , pages=1360–1371 , doi-access=free }
Corrigendum
/ref> {{cite journal , last=Austin , first=Jeremy J. , year=1996 , title=Molecular Phylogenetics of ''Puffinus'' Shearwaters: Preliminary Evidence from Mitochondrial Cytochrome ''b'' Gene Sequences , journal=Molecular Phylogenetics and Evolution , doi=10.1006/mpev.1996.0060 , pmid=8812308 , volume=6 , issue=1 , pages= 77–88 {{cite web, url=http://news.nationalgeographic.com/news/2006/08/060808-bird-migration.html, title=Longest Animal Migration Measured, Bird Flies 40,000 Miles a Year


External links


Shearwater videos
on the Internet Bird Collection * Seabirds Bird common names