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The minute black scavenger flies or "dung midges", are a family, Scatopsidae, of nematoceran flies. Despite being distributed throughout the world, they form a small family with only around 250 described species in 27 genera, although many await description and doubtless even more await discovery. These are generally small, sometimes minute, dark flies (from 0.6 to 5 mm), generally similar to black flies (Simuliidae), but usually lacking the humped thorax characteristic of that family. The larvae of most species are unknown, but the few that have been studied have a rather flattened shape and are terrestrial and saprophagous. Scatopsids are a well established group and fossils are known from amber deposits dating back to the Cretaceous
Cretaceous
period. Scatopse notata
Scatopse notata
(Linnaeus, 1758) is a cosmopolitan species. Its larval stages are found in decaying plant and animal material. Genera[edit] This list is incomplete; you can help by expanding it.

Anapausis Enderlein, 1912 Apiloscatopse Cook, 1874 Arthria Kirby, 1837 Aspistes Meigen, 1818 Austroclemina Borneoscatopse Brahemyia Amorim, 2007 Coboldia Melander, 1916 (sometimes erroneously as Colboldia) Colobostema Enderlein, 1926 Cooka Amorin, 2007 Diamphidicus Cook, 1971 Efcookella Ectaetia Enderlein, 1912 Ferneiella Cook, 1974 Hawomersleya Cook, 1971 Holoplagia Enderlein, 1912 † Mesoscatopse Neorhegmoclemina Parascatopse Cook, 1955 Parmaferia Cook, 1977 † Procolobostema † Protoscatopse Psectrosciara Kieffer, 1911 Quateiella Cook, 1975 Reichertella Enderlein, 1912 Rhegmoclema Enderlein, 1912 Rhegmoclemina Enderlein, 1936 Rhexoza Enderlein, 1936 Scatopse Geoffroy, 1762 (sometimes erroneously as Scatops or Scathops) Swammerdamella Enderlein, 1912 Thripomorpha Enderlein, 1905

Name[edit] The family name Scatopsidae
Scatopsidae
literally translates to "looks like feces" (from Greek skat "dung" and opsi "appearance"), but this seems to be a misinterpretation. It is derived from the genus Scatopse, which was misspelled as Scatops. See also[edit]

List of Australian Scatopsidae

References[edit]

[1] [2] - Family descriptions Australasian/Oceanian Diptera Catalog (Google cache)

Further reading[edit] Species descriptions[edit]

Cook, E.F. (1969). A synopsis of the Scatopsidae
Scatopsidae
of the Palaearctic Part I. Rhegmoclematini. Journal of Natural History
Journal of Natural History
3(3): 393-407 doi:10.1080/00222936900770341 (HTML abstract) Cook, E.F. (1972). A synopsis of the Scatopsidae
Scatopsidae
of the Palaearctic Part II. Swammerdamellini. Journal of Natural History
Journal of Natural History
6(6): 625-634. doi:10.1080/00222937200770561 Cook, E.F. (1974). A Synopsis of the Scatopsidae
Scatopsidae
of the palaearctic Part III. The Scatopsini. Journal of Natural History
Journal of Natural History
8(1): 61-100 doi:10.1080/00222937400770061

Fossil
Fossil
record[edit]

de Souza Amorim, D. (1998). Amber
Amber
Fossil
Fossil
Scatopsidae
Scatopsidae
(Diptera: Psychodomorpha). I. Considerations on Described Taxa, Procolobostema roseni, new species, from Dominican Amber, and the Position of Procolobostema in the Family. American Museum Novitates 3227; 1-17. PDF fulltext

External links[edit]

Scatopsidae
Scatopsidae
in Italian [https://www.biolib.cz/en/taxontree/id17379 BioLib: Incomplete taxonomic tree of Scatopsidae List of nearctic Scatopsidae
Scatopsidae
species Picture of Rhegmoclema hubachecki Diptera.info Images EOL images

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Extant Diptera families

Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera Superorder: Endopterygota

Suborder Nematocera

Axymyiomorpha

Axymyiidae

Culicomorpha

Culicoidea

Dixidae
Dixidae
(meniscus midges) Corethrellidae
Corethrellidae
(frog-biting midges) Chaoboridae
Chaoboridae
(phantom midges) Culicidae (mosquitoes)

Chironomoidea

Thaumaleidae
Thaumaleidae
(solitary midges) Simuliidae (black flies) Ceratopogonidae
Ceratopogonidae
(biting midges) Chironomidae
Chironomidae
(non-biting midges)

Blephariceromorpha

Blephariceridae
Blephariceridae
(net-winged midges) Deuterophlebiidae (mountain midges) Nymphomyiidae

Bibionomorpha

Bibionoidea

Bibionidae
Bibionidae
(march flies, lovebugs)

Anisopodoidea

Anisopodidae
Anisopodidae
(wood gnats)

Sciaroidea (fungus gnats)

Bolitophilidae Diadocidiidae Ditomyiidae Keroplatidae Mycetophilidae Sciaridae
Sciaridae
(dark-winged fungus gnats) Cecidomyiidae
Cecidomyiidae
(gall midges)

Psychodomorpha

Scatopsoidea

Canthyloscelidae Perissommatidae Scatopsidae
Scatopsidae
(minute black scavenger flies, or dung midges)

Psychodoidea

Psychodidae (moth flies)

Ptychopteromorpha

Ptychopteridae
Ptychopteridae
(phantom crane flies) Tanyderidae (primitive crane flies)

Tipulomorpha

Trichoceroidea

Trichoceridae
Trichoceridae
(winter crane flies)

Tipuloidea

Pediciidae
Pediciidae
(hairy-eyed craneflies) Tipulidae (crane flies)

Suborder Brachycera

Asilomorpha

Asiloidea

Apioceridae (flower-loving flies) Apsilocephalidae Apystomyiidae Asilidae
Asilidae
(robber flies) Bombyliidae
Bombyliidae
(bee flies) Evocoidae Hilarimorphidae (hilarimorphid flies) Mydidae (mydas flies) Mythicomyiidae Scenopinidae
Scenopinidae
(window flies) Therevidae
Therevidae
(stiletto flies)

Empidoidea

Atelestidae Hybotidae
Hybotidae
(dance flies) Dolichopodidae
Dolichopodidae
(long-legged flies) Empididae
Empididae
(dagger flies, balloon flies)

Nemestrinoidea

Acroceridae
Acroceridae
(small-headed flies) Nemestrinidae
Nemestrinidae
(tangle-veined flies)

Muscomorpha

Aschiza

Platypezoidea

Phoridae
Phoridae
(scuttle flies, coffin flies, humpbacked flies) Opetiidae
Opetiidae
(flat-footed flies) Ironomyiidae (ironic flies) Lonchopteridae
Lonchopteridae
(spear-winged flies) Platypezidae
Platypezidae
(flat-footed flies)

Syrphoidea

Syrphidae (hoverflies) Pipunculidae
Pipunculidae
(big-headed flies)

Schizophora

Acalyptratae

Conopoidea

Conopidae
Conopidae
(thick-headed flies)

Tephritoidea

Pallopteridae
Pallopteridae
(flutter flies) Piophilidae
Piophilidae
(cheese flies) Platystomatidae
Platystomatidae
(signal flies) Pyrgotidae Richardiidae Tephritidae
Tephritidae
(peacock flies) Ulidiidae
Ulidiidae
(picture-winged flies)

Nerioidea

Cypselosomatidae Micropezidae
Micropezidae
(stilt-legged flies) Neriidae
Neriidae
(cactus flies, banana stalk flies)

Diopsoidea

Diopsidae (stalk-eyed flies) Gobryidae Megamerinidae Nothybidae Psilidae
Psilidae
(rust flies) Somatiidae Strongylophthalmyiidae Syringogastridae Tanypezidae

Sciomyzoidea

Coelopidae
Coelopidae
(kelp flies) Dryomyzidae Helosciomyzidae Ropalomeridae Huttoninidae Heterocheilidae Phaeomyiidae Sepsidae
Sepsidae
(black scavenger flies) Sciomyzidae
Sciomyzidae
(marsh flies)

Sphaeroceroidea

Chyromyidae Heleomyzidae Sphaeroceridae
Sphaeroceridae
(small dung flies) Nannodastiidae

Lauxanioidea

Celyphidae
Celyphidae
(beetle-backed flies) Chamaemyiidae
Chamaemyiidae
(aphid flies) Lauxaniidae

Opomyzoidea

Agromyzidae
Agromyzidae
(leaf miner flies) Anthomyzidae Asteiidae Aulacigastridae (sap flies) Clusiidae
Clusiidae
(lekking, or druid flies) Fergusoninidae Marginidae Neminidae Neurochaetidae (upside-down flies) Odiniidae Opomyzidae Periscelididae Teratomyzidae Xenasteiidae

Ephydroidea

Camillidae Curtonotidae
Curtonotidae
(quasimodo flies) Diastatidae
Diastatidae
(bog flies) Ephydridae
Ephydridae
(shore flies) Drosophilidae
Drosophilidae
(vinegar and fruit flies)

Carnoidea

Acartophthalmidae Australimyzidae Braulidae
Braulidae
(bee lice) Canacidae
Canacidae
(beach flies) Carnidae Chloropidae
Chloropidae
(frit flies) Cryptochaetidae Inbiomyiidae Milichiidae
Milichiidae
(freeloader flies)

Lonchaeoidea

Cryptochetidae Lonchaeidae
Lonchaeidae
(lance flies)

Calyptratae

Muscoidea

Anthomyiidae
Anthomyiidae
(cabbage flies) Fanniidae
Fanniidae
(little house flies) Muscidae
Muscidae
(house flies, stable flies) Scathophagidae
Scathophagidae
(dung flies)

Oestroidea

Calliphoridae
Calliphoridae
(blow-flies: bluebottles, greenbottles) Mystacinobiidae (New Zealand batfly) Oestridae (botflies) Rhinophoridae Sarcophagidae (flesh flies) Tachinidae
Tachinidae
(tachina flies)

Hippoboscoidea

Glossinidae (tsetse flies) Hippoboscidae
Hippoboscidae
(louse flies) Mormotomyiidae
Mormotomyiidae
(frightful hairy fly) Nycteribiidae
Nycteribiidae
(bat flies) Streblidae
Streblidae
(bat flies)

Stratiomyomorpha

Stratiomyoidea

Pantophthalmidae
Pantophthalmidae
(timber flies) Stratiomyidae
Stratiomyidae
(soldier flies) Xylomyidae
Xylomyidae
(wood soldier flies)

Tabanomorpha

Rhagionoidea

Austroleptidae Bolbomyiidae Rhagionidae
Rhagionidae
(snipe flies)

Tabanoidea

Athericidae
Athericidae
(water snipe flies) Oreoleptidae Pelecorhynchidae Tabanidae (horse and deer flies)

Vermileonomorpha

Vermileonoidea

Vermileonidae

Xylophagomorpha

Xylophagoidea

Xylophagidae
Xylophagidae
(awl flies)

List of families of Diptera

Taxon identifiers

Wd: Q1265805 ADW: Scatopsidae BugGuide: 37474 EoL: 9005 EPPO: 1SCTSF Fauna Europaea: 11681 Fossilworks: 138435 GBIF: 6927 iNaturalist: 203230 ITIS: 125618 NCBI: 5

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