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''Santalum'' is a genus of woody
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the
Santalaceae The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial herbs, and epiphytic climbersHewson & George t al.br>''Santalaceae'' taxonomy, 1984, pp. 191-194.) which, like other mem ...
family, the best known and commercially valuable of which is the Indian
sandalwood Sandalwood is a class of woods from trees in the genus ''Santalum''. The woods are heavy, yellow, and fine-grained, and, unlike many other aromatic woods, they retain their fragrance for decades. Sandalwood oil is extracted from the woods for us ...
tree, '' S. album''. Members of the genus are
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s or shrubs. Most are root
parasites Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...
which
photosynthesize Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
their own food, but tap the roots of other species for water and inorganic
nutrients A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excre ...
. Several species, most notably ''S. album'', produce highly aromatic wood, used for scents and
perfume Perfume (, ; french: parfum) is a mixture of fragrant essential oils or aroma compounds (fragrances), fixatives and solvents, usually in liquid form, used to give the human body, animals, food, objects, and living-spaces an agreeable scent. Th ...
s and for herbal medicine. About 25 known species range across the
Indomalayan The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Indi ...
, Australasian, and
Oceanian realm The Oceanian realm is one of the World Wildlife Fund (WWF) biogeographic realms, and is unique in not including any continental land mass. It has the smallest land area of any of the WWF realms. This realm includes the islands of the Pacific Oc ...
s, from
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
through
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. Th ...
to the Pacific Islands, as far as
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state ...
and the
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
off the coast of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
. Indian sandalwood (''S. album'') is found in the tropical dry deciduous forests of India, the Lesser Sunda Islands of
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, and Arnhem Land of northern Australia. It is the only species of the genus found on the
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
n mainland, and may have been introduced to India from the Lesser Sundas centuries ago. Indian sandalwood has been stripped from most of India's forests, and is now rare in the wild. Five species, including ''S. album'', are native to Australia. '' S. acuminatum'', known as the sweet quandong or native peach, produces a shiny bright red fruit used increasingly in Australia for jams, jellies,
chutney A chutney is a spread in the cuisines of the Indian subcontinent. Chutneys are made in a wide variety of forms, such as a tomato relish, a ground peanut garnish, yogurt or curd, cucumber, spicy coconut, spicy onion or mint dipping sa ...
s, and pies. Four species, commonly called ''iliahi'', are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to Hawaii. '' S. fernandezianum'', endemic to the
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
off the coast of
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, was also
overexploited Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term ap ...
for its aromatic wood, and may now be extinct. ''Santalum'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e of some Lepidoptera species, including ''
Endoclita malabaricus ''Endoclita malabaricus'' is a species of moth of the family Hepialidae. It is known from India. Food plants for this species include ''Acacia'', ''Ailanthus'', ''Albizia'', ''Bridelia'', '' Cajanus'', ''Callicarpa'', ''Camellia'', '' Cassia'', ' ...
''.


Cultivation

The initial challenges to cultivating ''Santalum'' are numerous, not only due to its germination and growth needs, but also the amount of growing time required for the tree to properly mature. Germination of ''Santalum''
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s is not completely understood. Seeds cannot be effectively stored, and must be planted upon harvesting them from a fruiting tree. Even in doing this, the seeds may not germinate. As such, growing saplings can be quite labour-intensive. Furthermore, although ''Santalum'' trees
photosynthesize Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
on their own, the trees are hemiparasitic, with roots that seek out and tap the root systems of surrounding trees for water and nutrients. As such, each sapling is usually grown next to four or five host trees.
Pruning Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots. The practice entails the ''targeted'' removal of diseased, damaged, dead, ...
of host trees is also needed at times, since ''Santalum'' trees require much sunlight for growth. To produce commercially valuable sandalwood with high levels of fragrance oils, the trees have to be at least 40 years of age, but 80 or above is preferred. As such, those who begin cultivation of ''Santalum'' likely will not live to reap the rewards of their work. However, inferior sandalwood that has been cut or toppled at 30 years old can still fetch a decent price due to the demand for real sandalwood. Beyond these initial difficulties, growing sandalwood is not difficult, since it becomes more resistant to environmental stresses, pests, and diseases as it matures.


Species

* '' S. acuminatum'' A.DC. — desert quandong, sweet quandong, native peach ( Australia) * '' S. album'' L. — Indian sandalwood, white sandalwood, ''chandana'' (
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, northern Australia) * '' S. austrocaledonicum'' Vieill. ( New Caledonia,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
) * '' S. boninense'' (Nakai) Tuyama (
Bonin Islands The Bonin Islands, also known as the , are an archipelago of over 30 subtropical and tropical islands, some directly south of Tokyo, Japan and northwest of Guam. The name "Bonin Islands" comes from the Japanese word ''bunin'' (an archaic read ...
, Japan) * '' S. ellipticum'' Gaudich. — ''iliahialoe'', coast sandalwood (
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only state ...
) * '' S. fernandezianum'' Phil. (
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
) * '' S. freycinetianum'' Gaudich. — ''iliahi'' (Hawaii) * '' S. haleakalae'' Hillebr. — ''iliahi'' (Hawaii) * '' S. lanceolatum'' R.Br. — northern sandalwood (Australia) * '' S. macgregorii'' F.Muell (
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
) * '' S. murrayanum'' C.A.Gardner — bitter quandong (Australia) * '' S. obtusifolium'' (Australia) * '' S. paniculatum''
Hook. Sir William Jackson Hooker (6 July 178512 August 1865) was an English botanist and botanical illustrator, who became the first director of Kew when in 1841 it was recommended to be placed under state ownership as a botanic garden. At Kew h ...
& Arn.
— ''iliahi'' (Hawaii) * '' S. salicifolium'' — willowleaf sandalwood * '' S. spicatum'' (R.Br.) A.DC. — Australian sandalwood (Australia) * '' S. yasi''
Seem. Berthold Carl Seemann (25 February 1825, in Hanover, Germany – 10 October 1871, in Nicaragua, Central America), was a German botanist. He travelled widely and collected and described plants from the Pacific and South America. In 1844 he trave ...
- ''yasi'' ( Fiji,
Niue Niue (, ; niu, Niuē) is an island country in the South Pacific Ocean, northeast of New Zealand. Niue's land area is about and its population, predominantly Polynesian, was about 1,600 in 2016. Niue is located in a triangle between Tong ...
)
Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...
- ''Ahi''


References

* *


External links


Australian Quandong Industry Association




* * * {{Taxonbar, from=Q339027 Santalales genera