Salix hastata
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''Salix hastata'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
in the
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
, known by the common name halberd willow. It has an almost circumpolar distribution,''Salix hastata''.
The Nature Conservancy.
occurring throughout the northern
latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north ...
s of the Northern Hemisphere, most frequently found near the coast of the Arctic Ocean.Esser, Lora L. 1992
''Salix hastata''.
In: Fire Effects Information System, nline U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory.
In
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
, it occurs in the north and in the central mountains. It also occurs in northwestern Canada, and in Norway and Russia,''Salix hastata''.
Flora of North America.
as well as various alpine or mountainous areas of Eurasia.


Names

The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''hastata'' means "spear-shaped".


Description

This
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...
varies widely in appearance, depending on environmental conditions. In general, it is a
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...
growing 1 to 2 meters tall, but known to reach 4 meters. It forms
thicket A thicket is a very dense stand of trees or tall shrubs, often dominated by only one or a few species, to the exclusion of all others. They may be formed by species that shed large numbers of highly viable seeds that are able to germinate in t ...
s. The deciduous, alternately arranged leaves are up to 9.2 centimeters long. The leaves are often hairy, especially along the underside. The species is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
, with male and female reproductive parts occurring on separate plants. The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
is a
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
up to 5 centimeters long. The flowers are pollinated mainly by bees. The seeds
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
immediately upon contacting the substrate.


Habitat

Thickets of the shrub occur on slopes and riverbanks in the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, in
alpine climate Alpine climate is the typical weather (climate) for elevations above the tree line, where trees fail to grow due to cold. This climate is also referred to as a mountain climate or highland climate. Definition There are multiple definitions o ...
s, and on
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
. It is a
pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so ...
, colonizing recently cleared habitat, such as
floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...
s that have recently been scoured by water. It also grows in recently burned areas. Associated species include Barclay willow (''Salix barclayi''), Alaska willow (''Salix alaxensis''), tealeaf willow (''S. pulchra''), Richardson willow (''S. lanata'' ssp. ''richardsonii''),
black cottonwood ''Populus trichocarpa'', the black cottonwood, western balsam-poplar or California poplar, is a deciduous broadleaf tree species native to western North America. It is used for timber, and is notable as a model organism in plant biology. ...
(''Populus trichocarpa''),
alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...
(''Alnus'' spp.),
sedges The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
(''Carex'' ssp.), and
mosses Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornw ...
(''Polytrichum'' ssp.). On the
Alaska North Slope The Alaska North Slope ( Iñupiaq: ''Siḷaliñiq'') is the region of the U.S. state of Alaska located on the northern slope of the Brooks Range along the coast of two marginal seas of the Arctic Ocean, the Chukchi Sea being on the western sid ...
, ''S. hastata'' is a low-growing shrub. Sites that supported this and other low-growing willow species before being disturbed for construction of the
Trans-Alaska Pipeline System The Trans-Alaska Pipeline System (TAPS) is an oil transportation system spanning Alaska, including the trans-Alaska crude-oil pipeline, 11 pump stations, several hundred miles of feeder pipelines, and the Valdez Marine Terminal. TAPS is one of ...
were observed to have been recolonized by low-growing willows within four years after disturbance ceased. Natural regeneration of low-growing willows such as ''S. hastata'' was successful on moist riparian sites with silty soils, where they were mixed with the taller Alaska willow (''S. alaxensis''), and on dry sites with fine-textured soils. Halberd willow and Barclay willow are similar in appearance and distribution, and misidentifications have been reported in some cases in Alaska; the two species can be distinguished by the presence of sparse reddish hairs on the main leaf vein of Halberd willow. These two species of willow can also form natural hybrids with each other.


Varieties

This willow species has a few recognized
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
: *''S. hastata'' var. ''hastata'' L. *''S. hastata'' var. ''subintegrifolia'' Flod. *''S. hastata'' var. ''subalpina'' Anderss. *''S. hastata'' var. ''alpestris'' Anderss. ''S. hastata'' var. ''farriae'' is now considered a distinct species known as Farr's willow (''S. farriae'').


Ecology

Halberd willow is a food source for a number of mammals and birds. It is, however, not considered an important
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
for moose, as it is not as palatable compared to other willows such as Alaska willow, tealeaf willow, sandbar willow (''S. interior''), and littletree willow (''S. arbusculoides''). On the Alaska North Slope, low-growing willows such as Halberd, tealeaf, sandbar, and littletree willow are usually covered by snow during the winter, so they are not a winter food source for moose.


Cultivation

This species is cultivated in parks and gardens. The
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
'Wehrhahnii' has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


Other Uses

Native Americans used parts of willows, including this species, for medicinal purposes, basket weaving, to make bows and arrows, and for building animal traps. In
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
, willow leaves were chewed to treat mosquito bites and bee stings, as well as stomach aches.


References


External links

* {{Taxonbar, from=Q158425 hastata Plants described in 1753 Taxa named by Carl Linnaeus