The Russulales are an order of the Agaricomycetes, (which include the

agaric An agaric () is a type of fungus fruiting body characterized by the presence of a pileus (cap) that is clearly differentiated from the stipe (stalk), with lamellae (gills) on the underside of the pileus. In the UK, agarics are called "mushroom ...

genera ''

Russula ''Russula'' is a very large genus composed of around 750 worldwide species of ectomycorrhizal mushrooms. They are typically common, fairly large, and brightly colored – making them one of the most recognizable genera among mycologists and mushr ...

'' and '' Lactarius'' and their polyporoid and corticioid relatives). According to the ''Dictionary of the Fungi'' (10th edition, 2008), the order consists of 12

families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideal ...

, 80 genera, and 1767

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

. According to ''Species Fungorum'' (January 2016), the order contains 13 families, 117 genera (16 not assigned to a family), and 3,060 species. Russuloid agarics represent an independent evolutionary line of agarics, not directly related to the Agaricales. This group also includes a number of russuloid

hypogeous Hypogeal, hypogean, hypogeic and hypogeous (; ) are biological terms describing an organism's activity below the soil surface. In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain non ...

fungi,

polypores Polypores are a group of fungi that form large fruiting bodies with pores or tubes on the underside (see Delimitation for exceptions). They are a morphological group of basidiomycetes-like gilled mushrooms and hydnoid fungi, and not all polypo ...

such as ''

Bondarzewia ''Bondarzewia'' is a widely distributed genus of fungi in the family Bondarzewiaceae. The genus was circumscribed by mycologist Rolf Singer Rolf Singer (June 23, 1906 – January 18, 1994) was a German-born mycologist and one of the most im ...

'', some tooth fungi (e.g. '' Auriscalpium vulgare''), and

club fungi The clavarioid fungi are a group of fungi in the '' Basidiomycota'' typically having erect, simple or branched basidiocarps (fruit bodies) that are formed on the ground, on decaying vegetation, or on dead wood. They are colloquially called club fu ...

e.g. '' Artomyces''. Basidiospores in this group are typically ornamented with

amyloid Amyloids are aggregates of proteins characterised by a fibrillar morphology of 7–13 nm in diameter, a beta sheet (β-sheet) secondary structure (known as cross-β) and ability to be stained by particular dyes, such as Congo red. In the huma ...

warts or reticulation but a few exceptions are known, e.g. '' Heterobasidion annosum''. The genus ''

Clavicorona ''Clavicorona'' is a fungal genus in the family Auriscalpiaceae. The genus was first described by Maxwell Stanford Doty in 1947, who included the species ''C. pyxidata'', ''C. cristata'', ''C. taxophila'', and ''C. candelabrum''. ...

'' was often treated in the Russulales, but its

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

, '' C. taxophila'', is in the Agaricales. The remaining species are retained in the Russulales in the genus '' Artomyces''.

Genera ''Incertae sedis''

There are several genera classified in the Russulales that are i) poorly known, ii) have not been subjected to DNA analysis, or iii) if analysed phylogenetically do not group with as yet named or identified families, and have not been assigned to a specific family (i.e., ''

Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'' with respect to familial placement). These include: *'' Aleurocystidiellum'' *'' Dentipellopsis'' *'' Gloeoasterostroma'' *'' Gloeohypochnicium'' *''

Gloeopeniophorella ''Gloeopeniophorella'' is a genus of crust-like, wood-decaying fungi in the family Russulaceae. It contains six known species. ''Gloeopeniophorella'' was first described by Brazilian mycologist Johannes Rick Johannes (João Evangelista) Rick (1 ...

'' *'' Haloaleurodiscus'' *'' Neoalbatrellus'' *'' Polypus'' *'' Scopulodontia'' *'' Scytinostromella'' *'' Xeroceps''

References

External links

Russulales News

{{Taxonbar, from=Q219143 Basidiomycota orders