Roystonea
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''Roystonea'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of eleven
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of monoecious palms, native to the Caribbean Islands, and the adjacent coasts of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
(
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
),
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
.Kew World Checklist of Selected Plant Families
/ref> Commonly known as the royal palms, the genus was named after
Roy Stone Roy Stone (October 16, 1836 – August 5, 1905) was an American soldier, civil engineer, and inventor. He served in the American Civil War, distinguishing himself during the Battle of Gettysburg, and took part in the Spanish–American War. He ...
, a
U.S. Army The United States Army (USA) is the land service branch of the United States Armed Forces. It is one of the eight U.S. uniformed services, and is designated as the Army of the United States in the U.S. Constitution.Article II, section 2, cl ...
engineer Engineers, as practitioners of engineering, are professionals who invent, design, analyze, build and test machines, complex systems, structures, gadgets and materials to fulfill functional objectives and requirements while considering the limit ...
. It contains some of the most recognizable and commonly cultivated palms in tropical and subtropical regions.


Description

''Roystonea'' is a genus of large, unarmed, single-stemmed palms with
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves. The large stature and striking appearance of a ''Roystonea'' palm makes it a notable aspect of the landscape. The stems, which were compared to stone columns by
Louis Louis may refer to: * Louis (coin) * Louis (given name), origin and several individuals with this name * Louis (surname) * Louis (singer), Serbian singer * HMS ''Louis'', two ships of the Royal Navy See also Derived or associated terms * Lewis ( ...
and Elizabeth Agassiz in 1868, are smooth and columnar, although the trunks of '' R. altissima'' and '' R. maisiana'' are more slender than those of typical royal palms. Stems often are swollen and bulging along portions of their length, which may reflect years where growing conditions were better or worse than average. Leaf scars are often prominent along the stem, especially in young, rapidly growing individuals. Stem color ranges from gray-white to gray-brown except in '' R. violacea'', which have violet-brown or mauve stems. Royal palm, '' R. oleracea'', reaches heights of , but most species are in the range. The largest Royal palm is located in Floresta Estadual Edmundo Navarro de Andrade in
Rio Claro, São Paulo Rio Claro is a city in the state of São Paulo in Brazil. The elevation is 613 m. It was incorporated as the village of '' São João Batista do Ribeirão Claro'' in 1827, and this incorporation is celebrated every year on June 24 as a municipal h ...
, Brazil with 42.4 m and was discovered by Vincent Ferh and Mauro Galetti ''Roystonea'' leaves consist of a sheathing leaf base, a petiole, and a
rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...
. The leaf base forms a distinctive green sheath around the uppermost portion of the trunk. Known as the
crownshaft An elongated circumferential leaf base formation present on some species of palm is called a crownshaft. The leaf bases of some pinnate leaved palms (most notable being ''Roystonea regia'' or the royal palm but also including the genera ''Areca' ...
, this sheath extends down the trunk. The petiole connects the lead base with the rachis. Zona only reported petiole lengths for three of the 10 species, ranging from . The rachis is pinnately divided and ranges from long. The leaf segments themselves range in length from in ''R. altissima'' up to as much as in '' R. lenis''. They are arranged in two or three planes along the rachis. Many authors have reported that the leaves ''R. oleracea'' are arranged in a single plane, but American botanist Scott Zona reported that this is not the case. These plants have the ability to easily release their leaves in strong winds, a supposed adaption serving to prevent toppling during
hurricane A tropical cyclone is a rapidly rotating storm system characterized by a low-pressure center, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depend ...
s.
Inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
s occur beneath the crownshaft, emerging from a narrow, horn-shaped bract. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s on the branched panicles are usually white,
unisexual Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
, and contain both sexes. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is an oblong or globose drupe long and deep purple when ripe. Some species so closely resemble one another that scientific differentiation is by inflorescence detail; flower size, colour, etc.


Taxonomy

''Roystonea'' is placed in the subfamily Arecoideae and the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
Roystoneae, which only contains ''Roystonea''. The placement of ''Roystonea'' within the Arecoideae is uncertain; a
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...
based on plastid DNA failed to resolve the position of the genus within the Arecoideae. As of 2008, there appear to be no molecular phylogenetic studies of ''Roystonea''. One species is known only from two
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
ized flowers preserved in
Dominican amber Dominican amber is amber from the Dominican Republic derived from resin of the extinct tree '' Hymenaea protera''. Dominican amber differentiates itself from Baltic amber by being nearly always transparent, and it has a higher number of fossil inc ...
which were described in 2002.


Species

Accepted species:Carnevali, G., J. L. Tapia-Muñoz, R. Duno de Stefano & I. M. Ramírez Morillo. 2010. Flora Ilustrada de la Peninsula Yucatán: Listado Florístico 1–326


Distribution

''Roystonea'' has a circum-Caribbean distribution which ranges from southern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
in the north, to southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, Honduras and
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
in the east and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
and Colombia in the south. Species are found throughout the Caribbean, although only
Jamaica Jamaica (; ) is an island country situated in the Caribbean Sea. Spanning in area, it is the third-largest island of the Greater Antilles and the Caribbean (after Cuba and Hispaniola). Jamaica lies about south of Cuba, and west of His ...
and Hispaniola (with two native species) and
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
(with five native species) have more than one native species. A few species are planted in
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
as well.


Uses

Royal palms are widely planted for decorative purposes throughout their native region, and elsewhere in the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referr ...
and subtropics. They are considered by many to be the most beautiful palm in the world. Royal palms are very fond of water and thrive on supplemental
irrigation Irrigation (also referred to as watering) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has been devel ...
. They also do better in a soil with much humus. Though mainly a decorative plant, royal palms do have some minor agricultural uses. The heart of the palm is used to make salad in some parts of the Caribbean, and its seeds can be used as substitutes for coffee beans. Royal palm seeds were widely used in Cuba to feed pigs at least up to the 1940s and 1950s. The meat of pigs raised with royal palm seeds was said to be the very best. The lard obtained from pigs fattened or raised with royal palm seeds was said to exhibit a grainy texture, and by inference, to have been the best lard to consume. The seeds generally were obtained by men who specialized in climbing the royal palms using a set of two ropes looped around the stem, with two loops around the climber's legs to support himself. Once the climber reached the seed clumps, he would tie the clumps that were mature, cut them, and let them down by a rope supported from other seed clumps.


References


External links


Kew Palms Checklist: ''Roystonea''

Flora of North America: ''Roystonea''
* {{Taxonbar, from1=Q309496, from2=Q19450725 Arecaceae genera Neotropical realm flora National symbols of Haiti