The rough-toothed dolphin (''Steno bredanensis'') is a species of dolphin that can be found in deep warm and tropical waters around the world. The species was first described by

Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in na ...

in 1823. The genus name ''Steno'', of which this species is the only member, comes from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for 'narrow', referring to the animal's beak — which is a diagnostic characteristic of the species. The specific name honours van Breda, who studied Cuvier's writings. There are no recognised subspecies.

Taxonomy

This species was formerly classified in the now-defunct subfamily Stenoninae, but more recent evidence has found that, despite its prominent bill, it in fact belongs with the blunt-nosed dolphins in the subfamily

Globicephalinae Globicephalinae is a subfamily of oceanic dolphins that includes the pilot whales (''Globicephala'' spp.), the pygmy killer whale (''Feresa attenuata''), the rough-toothed dolphin (''Steno bredanensis''), the false killer whale (''Pseudorca cra ...

Description

The rough-toothed dolphin is a relatively large species, with adults ranging from in length, and weighing between ; males are larger than females. Its most visible characteristic feature is its conical head and slender nose; other dolphins either have a shorter snout or a more visibly bulging

melon A melon is any of various plants of the family Cucurbitaceae with sweet, edible, and fleshy fruit. The word "melon" can refer to either the plant or specifically to the fruit. Botanically, a melon is a kind of berry, specifically a "pepo". T ...

on the forehead. As the common name for the species implies, the teeth are also distinctive, having a roughened surface formed by numerous narrow irregular ridges. They have been reported to have between nineteen and twenty-eight teeth in each quarter of the jaw. The rough-toothed dolphins jaw on the outside is very distinct. The color of the lower jaw is usually white, but can have a hint of pink mixed in. The flippers are set back further along the body than in other similar dolphins, although, at sea this dolphin may be confused with

spinner Technology *Spinner (aeronautics), the aerodynamic cone at the hub of an aircraft propeller * Spinner (cell culture), laboratory equipment for cultivating plant or mammalian cells * Spinner (computing), a graphical widget in a GUI * Spinner (MIT Med ...

, spotted and

bottlenose dolphin Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the comm ...

s. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

is pronounced, being from in height. The animal's flanks are a light gray, while the back and dorsal fin are a much darker gray. Older individuals often have distinctive pinkish, yellow, or white markings around the mouth and along the underside.

Population and distribution

The distribution and population of the rough-toothed dolphin is poorly understood. They inhabit the

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

, and

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by ...

s, and in the

Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...

, in warm temperate to tropical waters, with occasional reports from cooler environments. Rough-toothed dolphins can also bee seen regularly in locations stretching from the

Windward Islands french: Îles du Vent , image_name = , image_caption = ''Political'' Windward Islands. Clockwise: Dominica, Martinique, Saint Lucia, Saint Vincent and the Grenadines, and Grenada. , image_alt = , locator_map = , location = Caribbean Sea No ...

Cape Verde , national_anthem = () , official_languages = Portuguese , national_languages = Cape Verdean Creole , capital = Praia , coordinates = , largest_city = capital , demonym ...

, but only a small handful have been seen in

Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...

and

Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...

. Live sightings are almost universally made far off-shore, beyond the

continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

, in water at least deep. Most of the research activity concerning the dolphin has been directed in the eastern Pacific, where a population estimate of 150,000 was obtained by researchers in the 1980s. Fossils belonging to the genus ''Steno'' are known from Europe and date to the early to mid-

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58 Rough-toothed dolphins are typically social animals, although solitary individuals are also sighted. An average group has between ten and twenty members, but they can vary from as few as two to as many as ninety. Such groups are thought to be temporary assemblages, composed of smaller, more permanent groups of two to eight closely related individuals that occasionally join together with others. They have also been reported to school together with other species of dolphin, and with

pilot whale Pilot whales are cetaceans belonging to the genus ''Globicephala''. The two extant species are the long-finned pilot whale (''G. melas'') and the short-finned pilot whale (''G. macrorhynchus''). The two are not readily distinguishable at sea, ...

false killer whale The false killer whale (''Pseudorca crassidens'') is a species of oceanic dolphin that is the only extant representative of the genus '' Pseudorca''. It is found in oceans worldwide but mainly in tropical regions. It was first described in 184 ...

s, and

humpback whale The humpback whale (''Megaptera novaeangliae'') is a species of baleen whale. It is a rorqual (a member of the family Balaenopteridae) and is the only species in the genus ''Megaptera''. Adults range in length from and weigh up to . The hu ...

s. Rough-toothed dolphins have been reported to bow-ride on a number of occasions, although apparently they do not do so as frequently as many other dolphin species. They do, however, commonly "skim", by swimming with their heads and chin above the surface of the water. They are known to be able to dive to at least and be able to stay underwater for at least fifteen minutes. Their echolocation clicks are unusually brief, lasting no more than 0.2 seconds, and have a relatively low frequency, ranging from 2.7 to 256 kHz, with a maximum peak frequency of 25 kHz. They also make longer whistles with a frequency between 3 and 12 kHz. Although details of their diet are sketchy, the stomach contents of stranded dolphins have included such fish such as silversides, sauries,

houndfish The houndfish (''Tylosurus crocodilus'') is a game fish of the family (biology), family Belonidae. It is the largest member of its family, growing up to in length and in weight. It is also often called the crocodile needlefish. Description Whi ...

smelts Smelts are a family of small fish, the Osmeridae, found in the North Atlantic and North Pacific Oceans, as well as rivers, streams and lakes in Europe, North America and Northeast Asia. They are also known as freshwater smelts or typical smelts ...

cutlassfish The cutlassfishes are about 45 species of predatory fish in the family Trichiuridae of the order Scombriformes found in seas throughout the world. Fish of this family are long, slender, and generally steely blue or silver in colour, giving ris ...

, and various

squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fittin ...

and

octopus An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, ...

es. Predators on rough-toothed dolphins are thought to include

killer whale The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus ''Orcinus'' and is recognizable by its black-and-white pat ...

s and

shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...

Reproduction

Rough-toothed dolphins give birth to a single young, after an unknown period of gestation; it is also unknown whether or not they have a distinct breeding season. The young are about long at birth, and grow rapidly for the first five years of life. Females reach sexual maturity somewhere between six and ten years of age, and males between five and ten years.

Conservation

The population is not believed to be threatened by human activities. A small number of individuals have been

harpoon A harpoon is a long spear-like instrument and tool used in fishing, whaling, sealing, and other marine hunting to catch and injure large fish or marine mammals such as seals and whales. It accomplishes this task by impaling the target ani ...

ed by Japanese whalers and pods are also slaughtered in the Taiji drive hunts. Others have been caught in seine nets by trawlers fishing for

tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...

. Less than a dozen rough-toothed dolphins live in dolphinaria around the world. The rough-toothed dolphin is covered by the Agreement on the Conservation of Small Cetaceans of the Baltic, North East Atlantic, Irish and North Seas (

ASCOBANS Agreement on the Conservation of Small Cetaceans of the Baltic, North East Atlantic, Irish and North Seas, often abbreviated to ASCOBANS, is a regional agreement on the protection of small cetaceans that was concluded as the Agreement on the Con ...

) and the Agreement on the Conservation of Cetaceans in the Black Sea, Mediterranean Sea and Contiguous Atlantic Area (

ACCOBAMS The Agreement on the Conservation of Cetaceans of the Black Sea, Mediterranean Sea and contiguous Atlantic Area, or ACCOBAMS, is a regional international treaty that binds its States Parties on the conservation of Cetacea in their territories. ...

). The species is further included in the Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia ( Western African Aquatic Mammals MoU) and the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region ( Pacific Cetaceans MoU). There are two rough toothed dolphins located at the

Clearwater Marine Aquarium Clearwater Marine Aquarium is a 501(c)(3) non-profit organization, and aquarium in Clearwater, Florida. It is dedicated to the rescue, rehabilitation and release of sick and injured marine animals, public education, conservation, and research. ...

. The facility is one of two that houses the species, in North America.

References

*''Steno bredanensis'' pp. 269–280, by J Maigret in ''Handbuch der Säugetiere Europas. Band 6: Meeressäuger Teil 1A: Wale und Delphine 1'' Niethammer J, Krapp F, (Eds.) (1995). {{Taxonbar , from=Q838812

rough-toothed dolphin The rough-toothed dolphin (''Steno bredanensis'') is a species of dolphin that can be found in deep warm and tropical waters around the world. The species was first described by Georges Cuvier in 1823. The genus name ''Steno'', of which this spe ...

Pantropical fauna Cosmopolitan mammals