Rock monitor
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The rock monitor (''Varanus albigularis'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of monitor lizard in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Varanidae The Varanidae are a family of lizards in the superfamily Varanoidea within the Anguimorpha group. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely rel ...
. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to Central,
East East or Orient is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fac ...
, and
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
. It is the second-longest lizard found on the continent, and the heaviest-bodied; locally, it is called ''leguaan'' or ''likkewaan''.


Taxonomy

First described by François Marie Daudin in 1802, ''V. albigularis'' has been classified as a subspecies of '' V. exanthematicus'', Laurent RF (1964). "A new subspecies of ''Varanus exanthematicus'' (Sauria, Varanidae)". ''Breviora'' (199): 1-5. (''Varanus exanthematicus ionidesii'', new subspecies). but has since been declared a distinct species based upon differences in hemipenal
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
. The
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name ''Varanus'' is derived from the
Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walter ...
word ''waral'' ورل, which is translated to English as "monitor". The specific name ''albigularis'' comes from a compound of two
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
words: ''albus'' meaning "white" and ''gula'' meaning "throat". The subspecies of ''V. albigularis'' are: * White-throated monitor, ''V. a. albigularis'' * Angolan white-throated monitor, ''V. a. angolensis'' *Eastern white-throated monitor, ''V. a. microstictus'' * Black-throated monitor, ''V. a. ionidesi'' "''Varanus albigularis'' ". The Reptile Database. www.reptile-database.org

/ref> (but may be synonymous with ''V. a. microstictus'')


Description

''Varanus albigularis'' is the heaviest-bodied lizard in Africa, as adult males average about and females weigh from . Large mature males can attain . It is the second longest African lizard after the
Nile monitor The Nile monitor (''Varanus niloticus'') is a large member of the monitor family (Varanidae) found throughout most of Sub-Saharan Africa and along the Nile, with invasive populations in North America. The population in West African forests and ...
(''Varanus niloticus''). ''Varanus albigularis'' reaches in total length (including tail), with its tail and body being of equal size. However, they rarely exceed 100-150cm in many areas. Mature specimens more typically will measure . The head and neck are the same length, and are distinct from each other. The bulbous, convex snout gives an angular, box-like appearance. The forked tongue is pink or bluish, and the body scales are usually a mottled gray-brown with yellowish or white markings.


Geographic range and habitat

''V. albigularis'' is found in
Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo ...
(
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
/
Zaire Zaire (, ), officially the Republic of Zaire (french: République du Zaïre, link=no, ), was a Congolese state from 1971 to 1997 in Central Africa that was previously and is now again known as the Democratic Republic of the Congo. Zaire was, ...
),
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
(
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label= Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalaha ...
, Republic of South Africa, Eswatini,
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and ...
,
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...
,
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
), the
African Great Lakes The African Great Lakes ( sw, Maziwa Makuu; rw, Ibiyaga bigari) are a series of lakes constituting the part of the Rift Valley lakes in and around the East African Rift. They include Lake Victoria, the second-largest fresh water lake in th ...
(
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
,
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
), and the Horn of Africa (
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
,
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
). ''V. albigularis'' is found in a variety of dry
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s, including steppes, prairies, and savannahs, but is absent from desert interiors, rainforests, and thick scrub forests.


Diet

''V. albigularis'' are generalists, feeding opportunistically on a broad variety of prey in the wild, such as other lizards, amphibians,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s,
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
s,
tortoise Tortoises () are reptiles of the family Testudinidae of the order Testudines (Latin: ''tortoise''). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like oth ...
s,
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s and small mammals. Tortoises make up a significant part of their diet, and are swallowed whole due to the hard shell. Otherwise, this species consumes very little vertebrate prey, eating primarily invertebrates, especially
millipedes Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
, beetles,
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estim ...
,
orthopterans Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshopp ...
and
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always en ...
s. Millipedes for example form nearly a quarter of their diet; the monitors are apparently resistant to its poisonous secretions. They are not averse to occasionally scavenging the corpses of vertebrate prey, even those as large as
vervet monkey The vervet monkey (''Chlorocebus pygerythrus''), or simply vervet, is an Old World monkey of the family Cercopithecidae native to Africa. The term "vervet" is also used to refer to all the members of the genus ''Chlorocebus''. The five distinct ...
s, which are sometimes torn to pieces by "death rolling" like a crocodilian prior to consumption. Live vertebrate prey other than tortoises are usually too fast to catch for these monitors, and therefore form very little of their diet. This contrasts with what is often a diet of mostly vertebrates in captivity, such as
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s,
poultry Poultry () are domesticated birds kept by humans for their eggs, their meat or their feathers. These birds are most typically members of the superorder Galloanserae (fowl), especially the order Galliformes (which includes chickens, quails, ...
or
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
.


Predator

Natural predators of adult rock monitors include
martial eagle The martial eagle (''Polemaetus bellicosus'') is a large eagle native to sub-Saharan Africa.Ferguson-Lees & Christie, ''Raptors of the World''. Houghton Mifflin Company (2001), . It is the only member of the genus ''Polemaetus''. A species of t ...
s and leopard.


Intelligence

An intelligent lizard, several specimens of ''V. albigularis'' have demonstrated the ability to count as high as six in an experiment conducted by Dr. John Philips at the
San Diego Zoo The San Diego Zoo is a zoo in Balboa Park, San Diego, California, housing 4000 animals of more than 650 species and subspecies on of Balboa Park leased from the City of San Diego. Its parent organization, San Diego Zoo Wildlife Alliance, is a p ...
in 1999. Philips offered varying numbers of
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class G ...
s, and the monitors were able to distinguish numbers whenever one was missing.King, Dennis; Green, Brian (1999). ''Goannas: The Biology of Varanid Lizards''. University of New South Wales Press. . p. 43.''The Weekend Australian''. July 24–25, 1999, p. 12.


Folklore

People living with the
HIV/AIDS Human immunodeficiency virus infection and acquired immunodeficiency syndrome (HIV/AIDS) is a spectrum of conditions caused by infection with the human immunodeficiency virus (HIV), a retrovirus. Following initial infection an individual ...
virus in
Yumbe District Yumbe District is a Districts of Uganda, district in Northern Region, Uganda. Like most other Ugandan districts, it is named after its 'chief town', Yumbe, where the district headquarters are located. Location Yumbe District is one of Uganda's mo ...
of
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
have been reported injecting themselves with the blood of rock monitors, which they believe to be a cure for the virus. Many are reportedly discontinuing
anti-retroviral therapy The management of HIV/AIDS normally includes the use of multiple antiretroviral drugs as a strategy to control HIV infection. There are several classes of antiretroviral agents that act on different stages of the HIV life-cycle. The use of multiple ...
to pursue this anecdotal treatment. As a result, ''V. albigularis'' is reported to have become an expensive item in the Ugandan black market, selling for more than 175 US$ each.


References


External links


Photo of ''Varanus albigularis''


Further reading

*Bayless, Mark K. (1992). "The Necropsy and internal Anatomy of a white-throated monitor lizard (''Varanus albigularis'' Daudin, 1802)". ''VaraNews'' 2 (1): 5-6. {{Taxonbar, from=Q935868 Varanus Reptiles described in 1802 Taxa named by François Marie Daudin