Reefs
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A reef is a ridge or
shoal In oceanography, geomorphology, and geoscience, a shoal is a natural submerged ridge, bank, or bar that consists of, or is covered by, sand or other unconsolidated material and rises from the bed of a body of water to near the surface. It ...
of rock,
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural,
abiotic In biology and ecology, abiotic components or abiotic factors are non-living chemical and physical parts of the environment that affect living organisms and the functioning of ecosystems. Abiotic factors and the phenomena associated with them under ...
processes—
deposition Deposition may refer to: * Deposition (law), taking testimony outside of court * Deposition (politics), the removal of a person of authority from political power * Deposition (university), a widespread initiation ritual for new students practiced f ...
of sand, wave erosion planing down rock outcrops, etc.—but there are also reefs such as the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s of tropical waters formed by biotic processes dominated by corals and
coralline algae Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of re ...
, and
artificial reef An artificial reef is a human-created underwater structure, typically built to promote marine life in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of trawling nets, or improve surfing. Many ...
s such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, to attract a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this. Earth's largest coral reef system is the
Great Barrier Reef The Great Barrier Reef is the world's largest coral reef system composed of over 2,900 individual reefs and 900 islands stretching for over over an area of approximately . The reef is located in the Coral Sea, off the coast of Queensland, A ...
in Australia, at a length of over .


Biotic

There is a variety of biotic reef types, including
oyster reef The term ''oyster reef'' refers to dense aggregations of oysters that form large colonial communities. Because oyster larvae need to settle on hard substrates, new oyster reefs may form on stone or other hard marine debris. Eventually the oyster ...
s and sponge reefs, but the most massive and widely distributed are tropical
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s. Although corals are major contributors to the framework and bulk material comprising a coral reef; the organisms most responsible for reef growth against the constant assault from ocean waves are
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an ad ...
algae, especially, although not entirely,
coralline algae Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of re ...
. The preferred substrate for oyster larvae is the shells of oysters so they tend to settle on adult oysters and thereby develop layers building upwards, eventually forming a fairly massive hard stony calcium carbonate structure on which other reef organisms like
sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
and seaweeds can grow, and provide a habitat for mobile benthic organisms. These biotic reef types take on additional names depending upon how the reef lies in relation to the land, if any. Reef types include
fringing reef A fringing reef is one of the three main types of coral reef. It is distinguished from the other main types, barrier reefs and atolls, in that it has either an entirely shallow backreef zone (lagoon) or none at all. If a fringing reef grows dire ...
s,
barrier reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. ...
s, and
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can gro ...
s. A fringing reef is a reef that is attached to an island. A barrier reef forms a calcareous barrier around an island, resulting in a
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into '' coastal lagoons'' (or ''barrier lagoons ...
between the shore and the reef. An atoll is a ring reef with no land present. The reef front (ocean side) is a high energy locale, whereas the internal lagoon will be at a lower energy with fine grained sediments.


Mounds

One definition distinguishes reefs from mounds as follows: both are considered to be varieties of organosedimentary buildups – sedimentary features, built by the interaction of organisms and their environment, that have synoptic relief and whose biotic composition differs from that found on and beneath the surrounding
sea floor The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean. All floors of the ocean are known as 'seabeds'. The structure of the seabed of the global ocean is governed by plate tectonics. Most of ...
. Reefs are held up by a macroscopic skeletal framework. Coral reefs are an example of this kind.
Corals Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
and calcareous algae grow on top of one another and form a three-dimensional framework that is modified in various ways by other organisms and inorganic processes. By contrast, mounds lack a macroscopic skeletal framework. Mounds are built by microorganisms or by organisms that don't grow a skeletal framework. A microbial mound might be built exclusively or primarily by
cyanobacteria Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, bl ...
. Examples of
biostrome A reef knoll is a land-based landform that comprises an immense pile of calcareous material that accumulated on a previously existing ancient sea floor. At the time of its accumulation it may have had enough structure from organisms such as sponges ...
s formed by cyanobacteria occur in the
Great Salt Lake The Great Salt Lake is the largest saltwater lake in the Western Hemisphere and the eighth-largest terminal lake in the world. It lies in the northern part of the U.S. state of Utah and has a substantial impact upon the local climate, particu ...
in
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
, United States, and in
Shark Bay Shark Bay ( Malgana: ''Gathaagudu'', "two waters") is a World Heritage Site in the Gascoyne region of Western Australia. The http://www.environment.gov.au/heritage/places/world/shark-bay area is located approximately north of Perth, on the ...
on the coast of
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to t ...
. Cyanobacteria do not have skeletons, and individuals are microscopic. Cyanobacteria can encourage the precipitation or accumulation of calcium carbonate to produce distinct sediment bodies in composition that have relief on the seafloor. Cyanobacterial mounds were most abundant before the evolution of shelly macroscopic organisms, but they still exist today;
stromatolite Stromatolites () or stromatoliths () are layered sedimentary formations ( microbialite) that are created mainly by photosynthetic microorganisms such as cyanobacteria, sulfate-reducing bacteria, and Pseudomonadota (formerly proteobacteria). T ...
s are microbial mounds with a laminated internal structure. Bryozoans and
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...
s, common contributors to marine sediments during the Mississippian, for instance, produced a very different kind of mound. Bryozoans are small and the skeletons of crinoids disintegrate. However, bryozoan and crinoid meadows can persist over time and produce compositionally distinct bodies of sediment with depositional relief. The
Proterozoic The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided i ...
Belt Supergroup The Belt Supergroup is an assemblage of primarily fine-grained sedimentary rocks and mafic intrusive rocks of late Precambrian (Mesoproterozoic) age. It is more than thick, covers an area of some 200,000 km2 (77,220 sq. mi), and is considered to ...
contains evidence of possible microbial mat and dome structures similar to stromatolite and chicken reef complexes.


Geologic

Rocky reefs are underwater outcrops of rock projecting above the adjacent unconsolidated surface with varying relief. They can be found in depth ranges from
intertidal The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
to deep water, and provide a substrate for a large range of sessile benthic organisms, and shelter for a large range of mobile organisms.


Structures

Ancient reefs buried within
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostra ...
sections are of considerable interest to
geologist A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, althou ...
s because they provide paleo-environmental information about the location in Earth's history. In addition, reef structures within a sequence of
sedimentary rock Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles ...
s provide a discontinuity which may serve as a trap or conduit for
fossil fuel A fossil fuel is a hydrocarbon-containing material formed naturally in the Earth's crust from the remains of dead plants and animals that is extracted and burned as a fuel. The main fossil fuels are coal, oil, and natural gas. Fossil fuels ma ...
s or mineralizing fluids to form
petroleum Petroleum, also known as crude oil, or simply oil, is a naturally occurring yellowish-black liquid mixture of mainly hydrocarbons, and is found in geological formations. The name ''petroleum'' covers both naturally occurring unprocessed crud ...
or ore deposits. Corals, including some major extinct groups
Rugosa The rugosa, also called the tetracorallia or horn coral, are an extinct order of solitary and colonial corals that were abundant in Middle Ordovician to Late Permian seas. Solitary rugosans (e.g., '' Caninia'', '' Lophophyllidium'', '' Neoza ...
and
Tabulata Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to ...
, have been important reef builders through much of the
Phanerozoic The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anim ...
since the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...
Period. However, other organism groups, such as calcifying algae, especially members of the red algae (
Rhodophyta Red algae, or Rhodophyta (, ; ), are one of the oldest groups of eukaryotic algae. The Rhodophyta also comprises one of the largest phyla of algae, containing over 7,000 currently recognized species with taxonomic revisions ongoing. The majorit ...
), and molluscs (especially the rudist bivalves during the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period) have created massive structures at various times. During the
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago ...
Period, the conical or tubular skeletons of
Archaeocyatha Archaeocyatha (or archaeocyathids 'ancient cups') is a taxon of extinct, sessile, reef-building marine sponges that lived in warm tropical and subtropical waters during the Cambrian Period. It is believed that the centre of the Archaeocyatha or ...
, an extinct group of uncertain affinities (possibly sponges), built reefs. Other groups, such as the Bryozoa, have been important interstitial organisms, living between the framework builders. The corals which build reefs today, the
Scleractinia Scleractinia, also called stony corals or hard corals, are marine animals in the phylum Cnidaria that build themselves a hard skeleton. The individual animals are known as polyps and have a cylindrical body crowned by an oral disc in which a ...
, arose after the
Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, ...
that wiped out the earlier rugose corals (as well as many other groups), and became increasingly important reef builders throughout the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
Era. They may have arisen from a rugose coral ancestor. Rugose corals built their skeletons of
calcite Calcite is a carbonate mineral and the most stable polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on the Mohs scale of mineral hardness, based on scratc ...
and have a different symmetry from that of the scleractinian corals, whose skeletons are
aragonite Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including pre ...
. However, there are some unusual examples of well-preserved aragonitic rugose corals in the
Late Permian Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
. In addition, calcite has been reported in the initial post-larval calcification in a few scleractinian corals. Nevertheless, scleractinian corals (which arose in the middle Triassic) may have arisen from a non-calcifying ancestor independent of the rugosan corals (which disappeared in the late Permian).


Artificial

An artificial reef is a human-created underwater structure, typically built to promote
marine life Marine life, sea life, or ocean life is the plants, animals and other organisms that live in the salt water of seas or oceans, or the brackish water of coastal estuaries. At a fundamental level, marine life affects the nature of the planet. ...
in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of
trawling Trawling is a method of fishing that involves pulling a fishing net through the water behind one or more boats. The net used for trawling is called a trawl. This principle requires netting bags which are towed through water to catch different spec ...
nets, or improve
surfing Surfing is a surface water sport in which an individual, a surfer (or two in tandem surfing), uses a board to ride on the forward section, or face, of a moving wave of water, which usually carries the surfer towards the shore. Waves suitable ...
. Many reefs are built using objects that were built for other purposes, for example by sinking oil rigs (through the Rigs-to-Reefs program), scuttling ships, or by deploying
rubble Rubble is broken stone, of irregular size, shape and texture; undressed especially as a filling-in. Rubble naturally found in the soil is known also as 'brash' (compare cornbrash)."Rubble" def. 2., "Brash n. 2. def. 1. ''Oxford English Dictionar ...
or construction debris. Other artificial reefs are purpose built (e.g. the reef balls) from PVC or concrete. Shipwrecks become artificial reefs on the seafloor. Regardless of construction method, artificial reefs generally provide stable hard surfaces where
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular micr ...
and invertebrates such as
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosiv ...
s, corals, and
oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not ...
s attach; the accumulation of attached marine life in turn provides intricate structure and food for assemblages of fish.


See also

* * * *


References


Sources

* Shears N.T. (2007) Biogeography, community structure and biological habitat types of subtidal reefs on the South Island West Coast, New Zealand. ''Science for Conservation 281''. p 53. Department of Conservation, New Zealand


External links


''Reef Rescue'' - Smithsonian Ocean Portal

Coral Reefs of the Tropics
: facts, photos and movies from
The Nature Conservancy The Nature Conservancy (TNC) is a global environmental organization headquartered in Arlington, Virginia. it works via affiliates or branches in 79 countries and territories, as well as across every state in the US. Founded in 1951, The Nat ...

NOAA Photo Library

Reef Environmental Education Foundation

NOS Data Explorer
- A portal to obtain NOAA National Ocean Service data


Atolls – Distribution, Development and Architecture
{{Authority control * Stratigraphy Coastal geography Coastal and oceanic landforms