Quercus coccinea
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''Quercus coccinea'', the scarlet oak, is a
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree in the red oak section ''Lobatae'' of the genus ''Quercus'', in the family Fagaceae. It is primarily distributed in the central and eastern United States. It occurs on dry, sandy, usually acidic soil. It is often an important canopy species in oak–heath forests. The scarlet oak is the official tree of Washington, D.C.


Description

''Quercus coccinea'' is a medium to large
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to around with an open, rounded crown; the maximum height is approximately . The trunk diameter at breast height is typically It is a medium-size tree that grows fast and matures relatively early. It sets a deep growing taproot. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are glossy green, long and broad, with seven lobes, and deep sinuses between the lobes. Each lobe has 3–7 bristle-tipped teeth. The leaf is hairless (unlike the related pin oak (''Q. palustris''), with tufts of pale orange-brown down where the lobe veins join the central vein). The foliage generally becomes bright scarlet in autumn. The flower color is yellow to green, depending on the season. The
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera '' Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and b ...
s are ovate, broad and long, a third to a half covered in a deep cup, green maturing pale brown about 18 months after
pollination Pollination is the transfer of pollen from an Stamen, anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by Anemophily, wind. Pollinating agents can ...
; the kernel is very bitter. Image:Quercus coccineum spring foliage and flower.jpg, Foliage and male flower in May, Exbury, UK File:2014-11-02 11 42 07 Scarlet Oak foliage during autumn along Lower Ferry Road in Ewing, New Jersey.JPG, Autumn foliage, Ewing, New Jersey


Similar species

It can be mistaken for the pin oak, the black oak (''Q. velutina''), or occasionally the red oak (''Q. rubra''). On the scarlet oak the sinuses between lobes are in comparison to pin oak, which has sinuses and the acorns are half covered by a deep cap. Additionally, pin oak foliage generally turns bronze in autumn.


Taxonomy

The common English name is derived from the fall (autumn) coloration of the foliage.


Distribution and habitat

It is primarily distributed in the central and eastern United States, from southern
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
west to
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
,
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and t ...
and
Missouri Missouri is a state in the Midwestern region of the United States. Ranking 21st in land area, it is bordered by eight states (tied for the most with Tennessee): Iowa to the north, Illinois, Kentucky and Tennessee to the east, Arkansas t ...
, and south as far as
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is bord ...
,
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
, and
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
. The tree needs full exposure to sunlight for best growth. Its preferred soil ranges from dry to somewhat dry well-drained, acidic soil.


Ecology


Forestry

April and May are flowering months for the scarlet oak. During this time, factors such as elevation and weather play a significant role in the flowering process. It takes about two seasons for the acorns to mature following maturation. Scarlet oak is prominent as a co-component of forests, including species such as white oak, black oak, and northern red oak. When at a lower elevation surrounding the Appalachian Mountains, pine forests and heaths are a common component. Oak seeds are faster-growing than many other trees and can compete very successfully. To regenerate oaks, the oak saplings should be 4–5 feet tall before removing the overstory. To favor oak regeneration, non-oak stems in the
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
exceeding 4 feet can be controlled by various methods. The quality of the environment impacts oak regeneration. Seed production starts when the tree is about 20 years old. Seed production reaches a maximum at approximately 50 years of age. The seed production (
masting Mast is the fruit of forest trees and shrubs, such as acorns and other nuts. The term derives from the Old English ''mæst'', meaning the nuts of forest trees that have accumulated on the ground, especially those used historically for fattening do ...
) is highly variable. A good masting year occurs every 3–5 years, depending on the weather and environment.


Wildlife value

Many species of wildlife look to the ''Q. coccinea'' for shelter, including small- to medium-sized birds, as well as small mammals such as squirrels. It is also a temporary home to various moth larvae throughout the year. It provides food in the form of acorns to many animals such as woodpeckers, blue jays, small mammals, wild turkeys, white-tailed deer, and black bears.


Threats

''Cryphonectria'' ''parasitica'', a bark pathogen, has been known to infect scarlet oak trees particularly in Pennsylvania. Cankers were found on many of these trees due to this parasite during 1989 and 1990. The parasite was removed from 69.7% of infected areas and recovered from 67% of incidents. Spongy moths are known to defoliate a scarlet oak, which can kill the tree. Other defoliating insects capable of killing scarlet oaks include the oak leaftier moth (''
Acleris semipurpurana ''Acleris semipurpurana'' is a species in the moth family Tortricidae, and one of several species of moth commonly known as oak leaftier or oak leaf tier. The larvae feed on the leaves of oak trees in the Eastern United States and southeastern ...
''), fall cankerworm ('' Alsophila pometaria''), forest tent caterpillar ('' Malacosoma disstria'')'','' and walking stick ('' Diapheromera femoral''). A few species of insects bore into the scarlet oak bark and trunk, including the towline chestnut borer ('' Agrilus bilineatus''), red oak borer ('' Enaphalodes rufulus''), oak timber worm ('' Arrenodes minutus''),
Ambrosia beetle Ambrosia beetles are beetles of the weevil subfamilies Scolytinae and Platypodinae ( Coleoptera, Curculionidae), which live in nutritional symbiosis with ambrosia fungi. The beetles excavate tunnels in dead, stressed, and healthy trees in wh ...
s (''
Scolytinae A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although the ...
'' and ''
Platypodinae Platypodinae is a weevil subfamily in the family Curculionidae. They are important early decomposers of dead woody plant material in wet tropics; all but two species are ambrosia beetles that cultivate fungi in tunnels excavated in dead wood as t ...
'' spp.), and the larvae of carpenterworms ('' Prionoxystus'' spp.). Gouty
galls Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to ...
are commonly found on smaller twigs and limbs due to the gouty oak
gall wasp Gall wasps, also incorrectly called gallflies, are hymenopterans of the family Cynipidae in the wasp superfamily Cynipoidea. Their common name comes from the galls they induce on plants for larval development. About 1,300 species of this gener ...
('' Callirhytis quercuspunctata''). The spongy oak apple gall wasp ('' Amphibolips confluenta'') causes gall growth on leaves and petioles. Black carpenter ants ('' Camponotus pennsylvanicus'') are known for nesting in the scarlet oak trees. 'Oak decline' can be caused by drought, moth defoliation, age, fire, environmental causes, and other factors. From 1968 to 1972, 27% of the scarlet oak population in the Newark watershed in New Jersey died from the spongy moth defoliation, which came before the towline chestnut borer and shoestring root rot attack.


Conservation status

Red oak is not listed in the CITES Appendices or on the IUCN Red List of Threatened Species.


Uses

Scarlet oak is sometimes planted as an
ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
, popular for its bright red fall color. The
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
'Splendens' has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
. Scarlet oak has an excellent red color during the fall months and is typically grown for shade and ornamental purposes. The red oak is a popular selection for lumber and is commonly used as a flooring material. Scarlet oak wood has a light to medium color consisting of reds and browns. The scarlet oak has a fairly coarse texture with a relatively large pore size. The durability of scarlet oak is less than the white oak, which has a higher level of decay and rot resistance. Scarlet oak lumber is easy to glue and looks attractive after staining and finishing. It has a distinct, appealing smell common with most oaks. Oak has been classified as a sensitizer, although it does not cause many allergic reactions. Reactions include eye and skin irritations and asthma-like symptoms, but not severe. White oak is typically more expensive than scarlet oak; red oak is in good supply with reasonable pricing, making it a very popular lumber product throughout the USA. Scarlet oak is commonly used in furniture, cabinetry, interior trim, flooring, and veneer. Scarlet oak shares many characteristics with red oak and often falls in the same category as red oak (''Quercus rubra'') in a broad sense


Medicinal

''Quercus coccinea'' produces galls from insect interactions which may be used to treat hemorrhages, chronic diarrhea, and dysentery.


References

*


External links


''Quercus coccinea'' images from Vanderbilt Unitersity Interactive Distribution Map of ''Quercus coccinea''photo of herbarium specimen at Missouri Botanical Garden, collected in Missouri in 1932
{{Taxonbar, from=Q854714 coccinea Endemic flora of the United States Hardwood forest plants Trees of the Eastern United States Trees of the Northeastern United States Trees of the Southeastern United States Trees of humid continental climate Flora of the Appalachian Mountains Natural history of the Great Smoky Mountains Plants described in 1770 Symbols of the District of Columbia Garden plants of North America Ornamental trees Taxa named by Otto von Münchhausen