Pyrus bourgaeana
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''Pyrus bourgaeana'', the Iberian pear, is a close relative of ''
Pyrus communis ''Pyrus communis'', the common pear, is a species of pear native to central and eastern Europe, and western Asia. It is one of the most important fruits of temperate regions, being the species from which most orchard pear cultivars grown in Eu ...
'' L. The latter was domesticated about 2500 years ago. This monoecious small tree (typically 3–6 m high) is widely distributed across the southern
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
and northern
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
, where it coexists with four ''Pyrus'' species: ''P. communis'' L., ''P. cordata'' Dew., ''P. spinosa'' Forssk, and ''P. nivalis'' Jacq. Characteristics to discriminate these species are the width of fruit peduncle, petal size, leaf width and petiole length served to the taxa. ''Pyrus bourgaeana'' flowers are white, rarely tinted pink, 2–3 cm diameter, and have five petals. Fruits are non- dehiscent
globose A sphere () is a geometrical object that is a three-dimensional analogue to a two-dimensional circle. A sphere is the set of points that are all at the same distance from a given point in three-dimensional space.. That given point is the ...
pomes weighing ~ 9.5 g, with green or brown skin inconspicuous to birds, copious lenticels permitting scent to emanate, and pulp high in fiber. Each fruit usually contains 2-4 full seeds. Within the Iberian Peninsula, ''P. bourgaeana'' distribution is very fragmented with trees occurring at low densities in small patches of
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western Europe, Western and Southern Europe and Anatolia, on the south by North Africa ...
scrubland that are isolated from each other by towns, cultivations, etc. Within these patches, mature trees often are aggregated in small clusters of 8-10 individuals Iberian pears are remarkably resistant to sicknesses or
blight Blight refers to a specific symptom affecting plants in response to infection by a pathogenic organism. Description Blight is a rapid and complete chlorosis, browning, then death of plant tissues such as leaves, branches, twigs, or floral org ...
; they are more often killed by storms or high winds than by sickness.


Phenology and interactions with animals

The Iberian pear flowers during February–March. It is pollinated by wide variety of insects such as bees, flies, and
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s. Each individual produces between 200-450 fruits that ripen and drop to the ground unaided from September to December. Predispersal seed losses by
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s (
microlepidoptera Microlepidoptera (micromoths) is an artificial (i.e., unranked and not monophyletic) grouping of moth families, commonly known as the 'smaller moths' (micro, Lepidoptera). These generally have wingspans of under 20 mm, and are thus harder to ...
larvae) are usually low. Its seeds most frequently are dispersed by mammals. Indeed, mammalian
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other s ...
s (
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by ...
s ''Meles meles'', red foxes ''Vulpes vulpes'', etc.) often ingest whole fruits fallen to the ground and disperse ingested seeds away from conspecifics with the capacity of germination. In some areas, however, these dispersers have declined to low densities by hunting and other human activities. Rabbits ''Oryctolagus cunniculus'' and some birds also feed on its fruit, eating part of the fruit pulp leaving the uneaten seeds under fruiting trees. Rodents often depredate the uncovered seeds in those fruits partially eaten by pulp feeders.
Ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...
s (
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of we ...
''Cervus elaphus'',
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
''Sus scrofa'') generally ingest whole fallen fruits but grind ingested seeds and mostly act as seed predators. Rodents also can act as post-dispersal predators of ''P. bourgaeana'' seeds. Seeds germinate epigeally either shortly after dispersal, or even within fallen fruits, and do not appear to persist in the soil seed bank. Seedlings emerge from winter to early spring, and extensive mortality occurs on young seedlings due to summer droughts and fungal infection.Fedriani, J. M., and M. Delibes. 2009a The strong aggregated patterning is thought to be the result of several non-exclusive processes. First, by creating the initial template on which post-dispersal processes act, its seed dispersers like foxes and badgers can be partially responsible for ''P. bourgaena'' aggregation. Second, dispersal limitation sometimes leads to seedling establishment beneath mother trees, resulting in an aggregated patterning. This is a likely possibility since a fraction of the fruit fallen beneath adult trees are not taken by mammals or are partially depulped by rabbits, without dispersing the seeds. Some seedlings emerge beneath mother trees every season and, eventually, a few of them get established. Finally, rhizome sprouting in response to disturbance could result in clustering if different sprouts emerge from a single individual and eventually produce fruit. In the
Doñana National Park Doñana National Park or Parque Nacional y Natural de Doñana is a natural reserve in Andalucía, southern Spain, in the provinces of Huelva (most of its territory), Cádiz and Seville. It covers , of which are a protected area. The park is an ...
(Southwestern Spain), ''P. bourgaeana'' experience heavy browsing by red deer ''Cervus elaphus'' and sprouts of a range of sizes emerge beneath some trees. As those shoots grow could eventually reach the adult size leading to tree clustering.


Uses and threats

Pear
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
(of any species) has one of the finest textures of the fruitwoods. It is prized for making
woodwind Woodwind instruments are a family of musical instruments within the greater category of wind instruments. Common examples include flute, clarinet, oboe, bassoon, and saxophone. There are two main types of woodwind instruments: flutes and re ...
instruments, and pear veneer is used in fine furniture. In southern Spain, the Iberian pear has been used as
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
for grafting pear
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s. The fruit is not edible for people. The major threats are urbanization and agricultural development.


References


External links

* {{Taxonbar, from=Q3285189 bourgaeana Flora of Morocco Flora of Spain Flora of Portugal Taxa named by Joseph Decaisne