Pucrasia
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The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
that lived during the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
. They are distantly related to pheasants and are most closely related to
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
and turkeys. Koklass are the only species in the monotypic genus ''Pucrasia''. Both the words ''koklass'' and ''pucrasia'' have been onomatopœically derived from the bird's territorial call. Koklass are boreal adapted species which separate into three distinct species groups. They are one of the few gamebirds that regularly fly uphill and are capable of sustained flights of many miles. They are monogamous with a slight tendency toward social
polyandry Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wives" ...
. Both parents rear the chicks. Koklass are largely vegetarian for much of the year consuming pine nuts, pine shoots, bamboo shoots and seeds. They are highly insectivorous during the warmer months that coincide with nesting and chick-rearing. During this phase of their life cycle they live almost exclusively on ants but also are documented consuming
catkins A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
,
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
and fruit. Koklass pheasant is a polytypic species, with nine
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
recognised so far. These are: # Indian koklass pheasant (''P. m. macrolopha''), # Western koklass pheasant (''P. m. castanea''), #
Kashmir koklass pheasant Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompass ...
(''P. m. biddulphi''), # Nepal koklass pheasant (''P. m. nipalensis''), # Meyer’s koklass pheasant (''P. m. meyeri''), # Orange-collared koklass pheasant (''P. m. ruficollis''), # Yellow-necked koklass pheasant (''P. m. xanthospila''), # Joret’s koklass pheasant (''P. m. jorentiana'') and # Darwin’s koklass pheasant (''P. m. darwini''). This entry deals with the subspecies ''P. m. biddulphi'', which ranges from
Kashmir Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompas ...
to Kullu in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
. With exception of the subspecies ''P. m. nipalensis'', ''P. m. castanea'' and ''P. m. macrolopha'', which are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the southern side of northwest and western
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
, the other five are confined to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
and
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
. The koklass pheasant is a medium-sized elusive bird confined to high altitude
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s from
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is borde ...
to central
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
, and in northeastern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
to northern and eastern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. Upper parts of male koklass pheasant are covered with silver-grey
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
streaked velvety-black down the centre of each feather, and it has the unique feature of a black head, chestnut breast and prominent white patches on the sides of neck. The females differ from males in above characters and instead their upper parts are covered with pale brown plumage. Both sexes, however, have distinct elongated tails tipped with pale feathers. The males are known to weigh about 1135–1415 g and the females, about 1025–1135 g, with the body length varying from 58–64 cm and 18–22 cm respectively. Immature and juveniles resemble adult females in plumage pattern. Like the
western tragopan The western tragopan or western horned tragopan (''Tragopan melanocephalus'') is a medium-sized brightly plumed pheasant found along the Himalayas from north-eastern districts of Khyber Pakhtunkhwa province in northern Pakistan in the west to Ut ...
, it does not extend its range above the tree line. One of the less colourful pheasants, the koklass exhibits moderate
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
. Though they skulk under bushes, which makes direct sighting difficult, they give loud chorus/predawn calls during the breeding season and during autumn, revealing their presence and allowing the populations to be estimated. They remain in pairs or small family groups throughout the year. They nest on the ground and spend the nights roosting on trees, or under rock overhangs.


Gallery

File:Koklass Pheasant @ Manila.jpg, ''Pucrasia macrolopha macrolopha'' Male at Manila,
Uttarakhand Uttarakhand ( , or ; , ), also known as Uttaranchal ( ; the official name until 2007), is a state in the northern part of India. It is often referred to as the "Devbhumi" (literally 'Land of the Gods') due to its religious significance and ...
File:Koklass Pheasant Manila3.jpg, ''Pucrasia macrolopha macrolopha'' Male at Manila,
Uttarakhand Uttarakhand ( , or ; , ), also known as Uttaranchal ( ; the official name until 2007), is a state in the northern part of India. It is often referred to as the "Devbhumi" (literally 'Land of the Gods') due to its religious significance and ...


References

{{Taxonbar, from=Q899810
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Ko ...
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Ko ...
Birds of Afghanistan Birds of China Birds of the Himalayas Birds of North India Birds of Nepal Birds of Pakistan
koklass pheasant The koklass pheasant (''Pucrasia macrolopha'') is a species of gamebird, being closely related to progenitive grouse that lived during the Miocene. They are distantly related to pheasants and are most closely related to grouse and turkeys. Ko ...
Taxa named by René Lesson