''Prenoceratops'', (meaning 'bent or prone-horned face' and derived from

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''prene-/πρηνη-'' meaning 'bent forwards' or 'prone', ''cerat-/κερατ-'' meaning 'horn' and ''-ops/ωψ'' meaning 'face') is a genus of

ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the Late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

. It was a relatively small dinosaur, reaching in length and in body mass. Its fossils have been found in the upper

Two Medicine Formation The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between and (million years ago), during Campanian (Late Cretaceous) time. It crops out to the east of the Rocky M ...

in the present-day U.S. state of

Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...

, in

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

age rock layers that have been dated to 74.3 million years ago.Ryan, M. J., Evans, D. C., Currie, P. J., Brown, C. M., & Brinkman, D. (2012). New leptoceratopsids from the Upper Cretaceous of Alberta, Canada. ''Cretaceous Research'', 35: 69-80. Fossils were also found in the

Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...

in the modern day

Canadian Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...

province of

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

, dating to around 77 million years ago.

Discovery and species

''Prenoceratops'' was first discovered on a

Blackfeet reservation The Blackfeet Nation ( bla, Aamsskáápipikani, script=Latn, ), officially named the Blackfeet Tribe of the Blackfeet Indian Reservation of Montana, is a federally recognized tribe of Siksikaitsitapi people with an Indian reservation in Monta ...

in Pondera County, Montana in layers coming from the

. The locality at which ''Prenoceratops'' was found was filled with many disassociated fossil elements of many individuals, known as a bonebed, though only the skull material has been described as of 2022. The holotype was one of the few associated specimens, including a surrangular fused with an articular, the specimen is designated as TCM 2003.1.1. ''Prenoceratops'' was later described by Brenda J. Chinnery in 2004, though the taxon has been little noticed since. It is unusual in that it is the only basal neoceratopsian known from a

bonebed A bone bed is any geological stratum or deposit that contains bones of whatever kind. Inevitably, such deposits are sedimentary in nature. Not a formal term, it tends to be used more to describe especially dense collections such as Lagerstätte. ...

and the sheer number of elements. An isolated right frontal from the

of southern Alberta, Canada was described in 2010 and ascribed to ''Prenoceratops'' as ''P.'' sp.Miyashita, T. E. T. S. U. T. O., Currie, P. J., Chinnery-Allgeier, B. J., Ryan, M. J., Chinnery-Allgeier, B. J., & Eberth, D. A. (2010). First basal neoceratopsian from the Oldman Formation (Belly River Group), southern Alberta. In ''New perspectives on horned dinosaurs: the Royal Tyrrell Museum Ceratopsian Symposium. Edited by MJ Ryan, BJ Chinnery-Allgeier, and DA Eberth. Indiana University Press, Bloomington, IN'' (pp. 83-90). The fossil was found near a fossilized nesting site of ''

Hypacrosaurus ''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high because it was almost but not quite as large as ''Tyrannosaurus'') was a genus of duckbill dinosaur similar in appearance to ''Co ...

''. Named ''Prenoceratops'' species include only ''P. pieganensis'' (type).

Classification

''Prenoceratops'' belonged to the Ceratopsia (which name is derived from Ancient Greek, meaning 'horned face'), a group of herbivorous dinosaurs with

parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoid ...

-like beaks, which thrived in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

during the Cretaceous Period. It is closely related to ''

Leptoceratops ''Leptoceratops'' (meaning 'Thin-horned face' and derived from Greek ''lepto-/λεπτο-'' meaning 'small', 'insignificant', 'slender', 'meagre' or 'lean', ''kerat-/κερατ-'' meaning 'horn' and ''-ops/ωψ'' meaning face), is a genus of lept ...

'', which it antedates by several million years. It is characterized by a lower, more sloping head than that of ''

''.

Diet

''Prenoceratops'', like all ceratopsians, was a

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

. During the Cretaceous, flowering plants were "geographically limited on the landscape", and so it is likely that this dinosaur fed on the predominant plants of the era:

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...

s and

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

s. It would have used its sharp ceratopsian beak to bite off the leaves or needles.

References

* * https://web.archive.org/web/20050225002043/http://www.vertpaleo.org/jvp/24-572-590.html (online abstract of the preceding article) * Leptoceratopsids Late Cretaceous dinosaurs of North America Fossil taxa described in 2004 Paleontology in Montana Campanian genus first appearances Campanian genus extinctions Ornithischian genera {{ceratopsian-stub

Discovery and species

Classification

Diet

See also

References