Polar Bear
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The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the
Arctic Circle The Arctic Circle is one of the two polar circles, and the most northerly of the five major circles of latitude as shown on maps of Earth. Its southern equivalent is the Antarctic Circle. The Arctic Circle marks the southernmost latitude at ...
, encompassing the
Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...
, its surrounding seas and surrounding land masses. It is the largest extant bear species, as well as the largest extant land carnivore. A boar (adult male) weighs around , while a sow (adult female) is about half that size. Although it is the sister species of the
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
, it has evolved to occupy a narrower
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...
, with many body characteristics adapted for cold temperatures, for moving across snow, ice and open water, and for hunting
seals Seals may refer to: * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impress an emblem, used as a means of a ...
, which make up most of its diet. Although most polar bears are born on land, they spend most of their time on the sea ice. Their scientific name means "
maritime Maritime may refer to: Geography * Maritime Alps, a mountain range in the southwestern part of the Alps * Maritime Region, a region in Togo * Maritime Southeast Asia * The Maritimes, the Canadian provinces of Nova Scotia, New Brunswick, and Pri ...
bear" and derives from this fact. Polar bears hunt their preferred food of seals from the edge of sea ice, often living off fat reserves when no sea ice is present. Because of their dependence on the sea ice, polar bears are classified as
marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their ...
s. Because of expected
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
caused by
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
, the polar bear is classified as a
vulnerable species A vulnerable species is a species which has been categorized by the International Union for Conservation of Nature as being threatened with extinction unless the circumstances that are threatening its survival and reproduction improve. Vulnera ...
. For decades, large-scale hunting raised international concern for the future of the species, but populations rebounded after controls and quotas began to take effect. For thousands of years, the polar bear has been a key figure in the material, spiritual, and cultural life of circumpolar peoples, and polar bears remain important in their cultures. Historically, the polar bear has also been known as the "white bear". It is sometimes referred to as the "nanook", based on the
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...
term ''nanuq''.


Naming and etymology

Constantine John Phipps Constantine John Phipps, 2nd Baron Mulgrave (30 May 1744 – 10 October 1792) was an English explorer and officer in the Royal Navy. He served during the Seven Years' War and the American War of Independence, seeing action in a number of b ...
was the first to describe the polar bear as a distinct species in 1774 in his report about his 1773 expedition towards the North Pole. He chose the scientific name ''Ursus maritimus'', the
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
for "maritime bear", due to the animal's native habitat. The
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...
refer to the animal as '' nanook'' ( transliterated as in the Inupiat language). The Yupik also refer to the bear as in Siberian Yupik. In the
Chukotko-Kamchatkan languages The Chukotko-Kamchatkan or Chukchi–Kamchatkan languages are a language family of extreme northeastern Siberia. Its speakers traditionally were indigenous hunter-gatherers and reindeer-herders. Chukotko-Kamchatkan is endangered. The Kamchatka ...
of Alyutor and Koryak, the name of the polar bear is , while in the related Chukchi, it is . In Russian, it is usually called (, 'white bear'), though an older word still in use is (, which comes from the Komi , "bear"). In
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirte ...
, the polar bear is referred to by the
french French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
terms ('white bear') or ('polar bear'). In Norwegian, one of the primary languages of the Svalbard archipelago, the polar bear is referred to as ('ice bear') or ('white bear'). The polar bear was previously considered to be in its own
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, ''Thalarctos''.This combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
words ''thalassa''/θαλασσα 'sea', and ''arctos''/αρκτος 'bear' and also, with reference to Ursa Major, 'northern' or 'of the north pole'
However, evidence of hybrids between polar bears and brown bears, and of the recent evolutionary divergence of the two species, does not support the establishment of this separate genus, and the accepted scientific name is now therefore ''Ursus maritimus'', as Phipps originally proposed.


Taxonomy and evolution

The bear
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
, Ursidae, is thought to have split from other carnivorans about 38 million years ago. The subfamily Ursinae originated approximately 4.2 million years ago. The oldest known polar bear
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
is a 130,000 to 110,000-year-old jaw bone, found on Prince Charles Foreland in 2004. Fossils show that between 10,000 and 20,000 years ago, the polar bear's
molar teeth The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone ...
changed significantly from those of the brown bear. Polar bears are thought to have diverged from a population of brown bears that became isolated during a period of glaciation in the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
from the eastern part of
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
(from Kamchatka and the Kolym Peninsula). The evidence from DNA analysis is more complex. The
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
(mtDNA) of the polar bear diverged from the brown bear, ''Ursus arctos'', roughly 150,000 years ago. Further, some
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
s of brown bear, as assessed by their mtDNA, were thought to be more closely related to polar bears than to other brown bears, meaning that the brown bear might not be considered a species under some
species concepts The species problem is the set of questions that arises when biologists attempt to define what a species is. Such a definition is called a species concept; there are at least 26 recognized species concepts. A species concept that works well for se ...
, but
paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ...
. The mtDNA of extinct Irish brown bears is particularly close to polar bears. A comparison of the nuclear genome of polar bears with that of brown bears revealed a different pattern, the two forming genetically distinct clades that diverged approximately 603,000 years ago, although the latest research is based on analysis of the complete genomes (rather than just the mitochondria or partial nuclear genomes) of polar and brown bears, and establishes the divergence of polar and brown bears at 400,000 years ago. However, the two species have mated intermittently for all that time, most likely coming into contact with each other during warming periods, when polar bears were driven onto land and brown bears migrated northward. Most brown bears have about 2 percent genetic material from polar bears, but one population, the ABC Islands bears, has between 5 percent and 10 percent polar bear genes, indicating more frequent and recent mating. Polar bears can breed with brown bears to produce fertile
grizzly–polar bear hybrid A grizzly–polar bear hybrid (also named grolar bear, pizzly bear, zebra bear, grizzlar, or nanulak) is a rare ursid hybrid that has occurred both in captivity and in the wild. In 2006, the occurrence of this hybrid in nature was confirmed by t ...
s; rather than indicating that they have only recently diverged, the new evidence suggests more frequent mating has continued over a longer period of time, and thus the two bears remain genetically similar. However, because neither species can survive long in the other's ecological niche, and because they have different morphology,
metabolism Metabolism (, from el, μεταβολή ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run ...
, social and feeding behaviours, and other
phenotypic In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological pr ...
characteristics, the two bears are generally classified as separate species. When the polar bear was originally documented, two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
were identified: the American polar bear (''Ursus maritimus maritimus'') by Constantine J. Phipps in 1774, and the Siberian polar bear (''Ursus maritimus marinus'') by
Peter Simon Pallas Peter Simon Pallas FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son of Professor of Surgery ...
in 1776. This distinction has since been invalidated. One alleged fossil subspecies has been identified: ''
Ursus maritimus tyrannus ''Ursus maritimus tyrannus'' () is a controversial extinct subspecies of polar bear. Discovery The subspecies was named by the Finnish paleontologist Björn Kurtén in 1964, based of a single fragmentary ulna found in the gravels of the Thames ...
'', which became extinct during the Pleistocene. ''U.m. tyrannus'' was significantly larger than the living subspecies. However, recent reanalysis of the fossil suggests that it was actually a brown bear.


Population and distribution

The polar bear is found in the Arctic Circle and adjacent land masses as far south as
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
. Due to the absence of human development in its remote habitat, it retains more of its original range than any other extant carnivore. While they are rare north of 88°, there is evidence that they range all the way across the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, and as far south as
James Bay James Bay (french: Baie James; cr, ᐐᓂᐯᒄ, Wînipekw, dirty water) is a large body of water located on the southern end of Hudson Bay in Canada. Both bodies of water extend from the Arctic Ocean, of which James Bay is the southernmost p ...
in Canada. Their southernmost range is near the boundary between the
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Genera ...
and
humid continental climate A humid continental climate is a climatic region defined by Russo-German climatologist Wladimir Köppen in 1900, typified by four distinct seasons and large seasonal temperature differences, with warm to hot (and often humid) summers and freez ...
zones. They can occasionally
drift Drift or Drifts may refer to: Geography * Drift or ford (crossing) of a river * Drift, Kentucky, unincorporated community in the United States * In Cornwall, England: ** Drift, Cornwall, village ** Drift Reservoir, associated with the village ...
widely with the sea ice, and there have been anecdotal sightings as far south as
Berlevåg ( sme, Bearalváhki) is a municipality in Troms og Finnmark county, Norway. It is located in the traditional district of Øst-Finnmark. The administrative centre of the municipality is the village of Berlevåg. There are two settlements in ...
on the Norwegian mainland and the
Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese language, Japanese: or ) are a volcanic archipelago currently administered as part of Sakh ...
in the Sea of Okhotsk. It is difficult to estimate a global population of polar bears as much of the range has been poorly studied; however, biologists use a working estimate of about 20–25,000 or 22–31,000 polar bears worldwide. With the discovery of a southeast Greenland population in 2022, there are 20 generally recognized, discrete subpopulations of polar bears. Aars, pp. 33–55. The subpopulations display seasonal fidelity to particular areas, but DNA studies show that they are not reproductively isolated. The 14 North American subpopulations range from the Beaufort Sea south to
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
and east to eastern
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
and account for about 54% of the global population. The usual range includes the territory of five nations:
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establish ...
(
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
),
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
( Svalbard),
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
, the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
(
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
) and
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
. These five nations are the signatories of the
International Agreement on the Conservation of Polar Bears The Agreement on the Conservation of Polar Bears is a multilateral treaty signed in Oslo, November 15, 1973, by the five nations with the largest polar bear populations: Canada, Denmark ( Greenland), Norway ( Svalbard), the United States, and the ...
, which mandates cooperation on research and conservation efforts throughout the polar bear's range. Bears sometimes swim to
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
from Greenland—about 600 sightings since the country's settlement in the 9th century, and five in the 21st century —and are always killed because of their danger, as well as the cost and difficulty of repatriation. Modern methods of tracking polar bear populations have been implemented only since the mid-1980s, and are expensive to perform consistently over a large area. The most accurate counts require flying a helicopter in the
Arctic climate The climate of the Arctic is characterized by long, cold winters and short, cool summers. There is a large amount of variability in climate across the Arctic, but all regions experience extremes of solar radiation in both summer and winter. Som ...
to find polar bears, shooting a tranquilizer dart at the bear to sedate it, and then tagging the bear. In
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
, some Inuit have reported increases in bear sightings around human settlements in recent years, leading to a belief that populations are increasing. Scientists have responded by noting that hungry bears may be congregating around human settlements, leading to the illusion that populations are higher than they actually are. The Polar Bear Specialist Group of the IUCN Species Survival Commission takes the position that "estimates of subpopulation size or sustainable harvest levels should not be made solely on the basis of traditional ecological knowledge without supporting scientific studies." Aars, pp. 61–62 Of the 19 polar bear subpopulations recognized in 2017, one was in decline, two were increasing, seven were stable, and nine had insufficient data.IUCN Polar Bear Specialist Group, 201
Summary of polar bear population status
.


Habitat

The polar bear is a
marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their ...
because it spends many months of the year at sea. However, it is the only living marine mammal with powerful, large limbs and feet that allow them to cover kilometres on foot and run on land. Its preferred habitat is the annual sea ice covering the waters over the
continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
and the Arctic inter-island
archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands, or sometimes a sea containing a small number of scattered islands. Examples of archipelagos include: the Indonesian Arch ...
s. These areas, known as the "Arctic ring of life", have high biological productivity in comparison to the deep waters of the high Arctic. The polar bear tends to frequent areas where sea ice meets water, such as polynyas and leads (temporary stretches of open water in Arctic ice), to hunt the seals that make up most of its diet. Freshwater is limited in these environments because it is either locked up in snow or saline. Polar bears are able to produce water through the metabolism of fats found in seal blubber, and are therefore found primarily along the perimeter of the polar ice pack, rather than in the Polar Basin close to the
North Pole The North Pole, also known as the Geographic North Pole or Terrestrial North Pole, is the point in the Northern Hemisphere where the Earth's axis of rotation meets its surface. It is called the True North Pole to distinguish from the Ma ...
where the density of seals is low.
Matthews Matthews may refer to: People * Matthews (surname) Places * Matthews Island, Antarctica * Matthews Range, Kenya * Mount Matthews, New Zealand United States * Matthews, Georgia * Matthews, Indiana * Matthews, Maryland * Matthews, Missouri * Mat ...
, p. 15
Annual ice contains areas of water that appear and disappear throughout the year as the weather changes. Seals migrate in response to these changes, and polar bears must follow their prey. In
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
,
James Bay James Bay (french: Baie James; cr, ᐐᓂᐯᒄ, Wînipekw, dirty water) is a large body of water located on the southern end of Hudson Bay in Canada. Both bodies of water extend from the Arctic Ocean, of which James Bay is the southernmost p ...
, and some other areas, the ice melts completely each summer (an event often referred to as "ice-floe breakup"), forcing polar bears to go onto land and wait through the months until the next freeze-up. In the Chukchi and Beaufort seas, most polar bears retreat each summer to the ice further north that remains frozen year-round, though a large portion of the population (15–40%) has been observed spending all summer on land every year in these areas since the 1980s.


Physical characteristics

The only other bear of comparable size to the polar bear is the Kodiak bear, which is a subspecies of
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
. Adult male polar bears weigh and measure in total length. Hemstock, p. 4 Around the Beaufort Sea, however, mature males reportedly average . Adult females are roughly half the size of males and normally weigh , measuring in length. Elsewhere, a slightly larger estimated average weight of was claimed for adult females. When pregnant, however, females can weigh as much as . The polar bear is among the most sexually dimorphic of mammals, surpassed only by the
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the ...
s such as elephant seals. The largest polar bear on record, reportedly weighing , was a male shot at Kotzebue Sound in northwestern Alaska in 1960. This specimen, when mounted, stood tall on its hindlegs. The shoulder height of an adult polar bear is . While all bears are short-tailed, the polar bear's tail is relatively the shortest amongst living bears, ranging from in length. Compared with its closest relative, the brown bear, the polar bear has a more elongated body build and a longer skull and nose. As predicted by Allen's rule for a northerly animal, the legs are stocky and the ears and tail are small. However, the feet are very large to distribute load when walking on snow or thin ice and to provide propulsion when swimming; they may measure across in an adult. Lockwood, pp. 10–16 The pads of the paws are covered with small, soft papillae (dermal bumps), which provide traction on the ice. The polar bear's claws are short and stocky compared to those of the brown bear, perhaps to serve the former's need to grip heavy prey and ice. The claws are deeply scooped on the underside which assists in digging in the ice of the natural habitat. Research of injury patterns in polar bear forelimbs found injuries to the right forelimb to be more frequent than those to the left, suggesting, perhaps, right-handedness. Unlike the brown bear, polar bears in captivity are rarely overweight or particularly large, possibly as a reaction to the warm conditions of most zoos. The 42 teeth of a polar bear reflect its highly carnivorous diet. The cheek teeth are smaller and more jagged than in the brown bear, and the canines are larger and sharper. The dental formula is . Polar bears are superbly insulated by up to of adipose tissue, their hide and their fur. Polar bear fur consists of a layer of dense underfur and an outer layer of guard hairs, which appear white to tan but are actually transparent. Two genes that are known to influence melanin production, LYST and AIM1, are both mutated in polar bears, possibly leading to the absence on this pigment in their fur. The guard hair is over most of the body. Polar bears gradually moult from May to August, but, unlike other Arctic mammals, they do not shed their coat for a darker shade to provide camouflage in summer conditions. The hollow guard hairs of a polar bear coat were once thought to act as fiber-optic tubes to conduct light to its black skin, where it could be absorbed; however, this hypothesis was disproved by a study in 1998. The white coat usually yellows with age. When kept in captivity in warm, humid conditions, the fur may turn a pale shade of green due to
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular micr ...
growing inside the guard hairs. Males have significantly longer hairs on their forelegs, which increase in length until the bear reaches 14 years of age. The male's ornamental foreleg hair is thought to attract females, serving a similar function to the
lion's mane Lion's mane may refer to: *Mane (lion), the mane of the adult male lion, the fur around its face *Lion's mane jellyfish * Lion's mane mushroom * Lion's mane nudibranch *" The Adventure of the Lion's Mane", a Sherlock Holmes short story by Sir Arthur ...
. The polar bear has an extremely well developed sense of smell, being able to detect seals nearly away and buried under of snow. Its hearing is about as acute as that of a human, and its vision is also good at long distances. Rosing, pp. 20–23 The polar bear is an excellent swimmer and often will swim for days. One bear swam continuously for 9 days in the frigid Bering Sea for to reach ice far from land. She then travelled another . During the swim, the bear lost 22% of her body mass and her yearling cub died. With its body fat providing buoyancy, the bear swims in a
dog paddle The dog paddle or doggy paddle is a simple swimming style. It is characterized by the swimmer lying on their chest and moving their hands and legs alternately in a manner reminiscent of how dogs and other quadrupedal mammals swim. It is effectiv ...
fashion using its large forepaws for propulsion. Polar bears can swim at . When walking, the polar bear tends to have a lumbering gait and maintains an average speed of around . When sprinting, they can reach up to .


Life history and behaviour

Unlike brown bears, polar bears are not
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
. Although stereotyped as being voraciously aggressive, they are normally cautious in confrontations, and often choose to escape rather than fight.
Matthews Matthews may refer to: People * Matthews (surname) Places * Matthews Island, Antarctica * Matthews Range, Kenya * Mount Matthews, New Zealand United States * Matthews, Georgia * Matthews, Indiana * Matthews, Maryland * Matthews, Missouri * Mat ...
, pp. 27–29
Satiated polar bears rarely attack humans unless severely provoked. However, due to their lack of prior human interaction, hungry polar bears are extremely unpredictable, fearless towards people and are known to kill and sometimes eat humans. Bruemmer, pp. 25–33 Many attacks by
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
s are the result of surprising the animal, which is not the case with the polar bear. Polar bears are stealth hunters, and the victim is often unaware of the bear's presence until the attack is underway. Whereas brown bears often maul a person and then leave, polar bear attacks are more likely to be predatory and are almost always fatal. However, due to the very small human population around the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, such attacks are rare. Michio Hoshino, a Japanese wildlife photographer, was once pursued briefly by a hungry male polar bear in northern Alaska. According to Hoshino, the bear started running but Hoshino made it to his truck. The bear was able to reach the truck and tore one of the doors off the truck before Hoshino was able to drive off. In general, adult polar bears live solitary lives. Yet, they have often been seen playing together for hours at a time and even sleeping in an embrace, and polar bear zoologist Nikita Ovsianikov has described adult males as having "well-developed friendships." Cubs are especially playful as well. Among young males in particular, play-fighting may be a means of practicing for serious competition during mating seasons later in life. Polar bears are usually quiet but do communicate with various sounds and vocalizations. Females communicate with their young with moans and chuffs, and the distress calls of both cubs and subadults consists of bleats. Cubs may hum while nursing. When nervous, bears produce huffs, chuffs and snorts while hisses, growls and roars are signs of aggression. Chemical communication can also be important: bears leave behind their scent in their tracks which allow individuals to keep track of one another in the vast Arctic wilderness. In 1992, a photographer near Churchill took a now widely circulated set of photographs of a polar bear playing with a
Canadian Eskimo Dog The Canadian Eskimo Dog or Canadian Inuit Dog is a breed of working dog from the Arctic. Other names include ''qimmiq'' The pair wrestled harmlessly together each afternoon for 10 days in a row for no apparent reason, although the bear may have been trying to demonstrate its friendliness in the hope of sharing the kennel's food. This kind of social interaction is uncommon; it is far more typical for polar bears to behave aggressively towards dogs. Rosing, pp. 128–132


Hunting and diet

The polar bear is the most carnivorous member of the bear family, and throughout most of its range, its diet primarily consists of ringed (''Pusa hispida'') and bearded seals (''Erignathus barbatus''). The Arctic is home to millions of seals, which become prey when they surface in holes in the ice in order to breathe, or when they haul out on the ice to rest.
Matthews Matthews may refer to: People * Matthews (surname) Places * Matthews Island, Antarctica * Matthews Range, Kenya * Mount Matthews, New Zealand United States * Matthews, Georgia * Matthews, Indiana * Matthews, Maryland * Matthews, Missouri * Mat ...
, pp. 73–88
Polar bears hunt primarily at the interface between ice, water, and air; they only rarely catch seals on land or in open water. The polar bear's most common hunting method is called ''still-hunting'': Hemstock, pp. 24–27 the bear uses its excellent sense of smell to locate a seal breathing hole, and crouches nearby in silence for a seal to appear. The bear may lie in wait for several hours. When the seal exhales, the bear smells its breath, reaches into the hole with a forepaw, and drags it out onto the ice. The polar bear kills the seal by biting its head to crush its skull. The polar bear also hunts by stalking seals resting on the ice: upon spotting a seal, it walks to within , and then crouches. If the seal does not notice, the bear creeps to within of the seal and then suddenly rushes forth to attack. A third hunting method is to raid the birth lairs that female seals create in the snow. A widespread legend tells that polar bears cover their black noses with their paws when hunting. This behaviour, if it happens, is rare – although the story exists in the oral history of northern peoples and in accounts by early Arctic explorers, there is no record of an eyewitness account of the behaviour in recent decades. Mature bears tend to eat only the calorie-rich skin and blubber of the seal, which are highly digestible, whereas younger bears consume the protein-rich red meat. Studies have also photographed polar bears scaling near-vertical cliffs, to eat birds' chicks and eggs. For subadult bears, which are independent of their mother but have not yet gained enough experience and body size to successfully hunt seals, scavenging the carcasses from other bears' kills is an important source of nutrition. Subadults may also be forced to accept a half-eaten carcass if they kill a seal but cannot defend it from larger polar bears. After feeding, polar bears wash themselves with water or snow. Although polar bears are extraordinarily powerful, its primary prey species, the ringed seal, is much smaller than itself, and many of the seals hunted are pups rather than adults. Ringed seals are born weighing and grown to an estimated average weight of only . They also in places prey heavily upon the harp seal (''Pagophilus groenlandicus''), the harbour seal (''Phoca vitulina''), or the hooded seal (''Cystophora cristata''). The bearded seal, on the other hand, can be nearly the same size as the bear itself, averaging . Adult male bearded seals, at are too large for a female bear to overtake, and so are potential prey only for mature male bears. Large males also occasionally attempt to hunt and kill even larger prey items. It can kill an adult walrus (''Odobenus rosmarus''), although this is rarely attempted. At up to and a typical adult mass range of , a walrus can be more than twice the bear's weight, has extremely thick skin and has up to -long ivory tusks that can be used as formidable weapons. A polar bear may charge a group of walruses, with the goal of separating a young, infirm, or injured walrus from the pod. They will even attack adult walruses when their diving holes have frozen over or intercept them before they can get back to the diving hole in the ice. Yet, polar bears will very seldom attack full-grown adult walruses, with the largest male walrus probably invulnerable unless otherwise injured or incapacitated. Since an attack on a walrus tends to be an extremely protracted and exhausting venture, bears have been known to back down from the attack after making the initial injury to the walrus. Polar bears have also been seen to prey on
beluga whale The beluga whale () (''Delphinapterus leucas'') is an Arctic and sub-Arctic cetacean. It is one of two members of the family Monodontidae, along with the narwhal, and the only member of the genus ''Delphinapterus''. It is also known as the ...
s (''Delphinapterus leucas'') and narwhals (''Monodon monoceros''), by swiping at them at breathing holes. The whales are of similar size to the walrus and nearly as difficult for the bear to subdue. Most terrestrial animals in the Arctic can outrun the polar bear on land as polar bears overheat quickly, and most marine animals the bear encounters can outswim it. In some areas, the polar bear's diet is supplemented by walrus calves and by the carcasses of dead adult walruses or whales, whose blubber is readily devoured even when rotten. Polar bears sometimes swim underwater to catch fish like the Arctic charr or the fourhorn sculpin. With the exception of pregnant females, polar bears are active year-round, although they have a vestigial hibernation induction trigger in their blood. Unlike brown and black bears, polar bears are capable of
fasting Fasting is the abstention from eating and sometimes drinking. From a purely physiological context, "fasting" may refer to the metabolic status of a person who has not eaten overnight (see " Breakfast"), or to the metabolic state achieved after ...
for up to several months during late summer and early fall, when they cannot hunt for seals because the sea is unfrozen. When sea ice is unavailable during summer and early autumn, some populations live off fat reserves for months at a time, as polar bears do not 'hibernate' any time of the year. Being both curious animals and scavengers, polar bears investigate and consume garbage where they come into contact with humans. Polar bears may attempt to consume almost anything they can find, including hazardous substances such as styrofoam,
plastic Plastics are a wide range of synthetic or semi-synthetic materials that use polymers as a main ingredient. Their plasticity makes it possible for plastics to be moulded, extruded or pressed into solid objects of various shapes. This adapta ...
, car batteries,
ethylene glycol Ethylene glycol ( IUPAC name: ethane-1,2-diol) is an organic compound (a vicinal diol) with the formula . It is mainly used for two purposes, as a raw material in the manufacture of polyester fibers and for antifreeze formulations. It is an o ...
, hydraulic fluid, and motor oil. The
dump Dump generally refers to a place for disposal of solid waste, a rubbish dump, or landfill. The word has other uses alone or in combination, and may refer to: * Midden, historically a dump for domestic waste * Dump job, a term for criminal disposal ...
in
Churchill Sir Winston Leonard Spencer Churchill (30 November 187424 January 1965) was a British statesman, soldier, and writer who served as Prime Minister of the United Kingdom twice, from 1940 to 1945 during the Second World War, and again from 1 ...
,
Manitoba Manitoba ( ) is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population o ...
was closed in 2006 to protect bears, and waste is now recycled or transported to Thompson, Manitoba.


Dietary flexibility

Although seal predation is the primary and an indispensable way of life for most polar bears, when alternatives are present they are quite flexible. Polar bears consume a wide variety of other wild foods, including muskox (''Ovibos moschatus''),
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subs ...
(''Rangifer tarandus''), birds, eggs,
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s,
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...
s, other
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s,
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
and other polar bears. They may also eat plants, including
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, ras ...
,
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the su ...
s, and kelp; however, none of these have been a significant part of their diet, except for beachcast
marine mammal Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their ...
carcasses. Given the change in climate, with ice breaking up in areas such as the
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
earlier than it used to, polar bears are exploiting food resources such as snow geese and eggs, and plants such as
lyme grass ''Leymus arenarius'' is a psammophilic ( sand-loving) species of grass in the family Poaceae, native to the coasts of Atlantic and Northern Europe. ''Leymus arenarius'' is commonly known as sand ryegrass, sea lyme grass, or simply lyme grass.< ...
in increased quantities. When stalking land animals, such as muskox,
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subs ...
, and even willow ptarmigan (''Lagopus lagopus''), polar bears appear to make use of vegetative cover and wind direction to bring them as close to their prey as possible before attacking. Polar bears have been observed to hunt the small Svalbard reindeer (''R. t. platyrhynchus''), which weigh only as adults, as well as the barren-ground caribou (''R. t. groenlandicus''), which is about twice as heavy as the former. Adult muskox, which can weigh or more, are a more formidable quarry. Although ungulates are not typical prey, the killing of one during the summer months can greatly increase the odds of survival during that lean period. Like the brown bear, most ungulate prey of polar bears is likely to be young, sickly or injured specimens rather than healthy adults. The polar bear's
metabolism Metabolism (, from el, μεταβολή ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run ...
is specialized to require large amounts of fat from marine mammals, and it cannot derive sufficient caloric intake from terrestrial food. In their southern range, especially near
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
and
James Bay James Bay (french: Baie James; cr, ᐐᓂᐯᒄ, Wînipekw, dirty water) is a large body of water located on the southern end of Hudson Bay in Canada. Both bodies of water extend from the Arctic Ocean, of which James Bay is the southernmost p ...
, Canadian polar bears endure all summer without sea ice to hunt from. Here, their food ecology shows their dietary flexibility. They still manage to consume some seals, but they are food-deprived in summer as only marine mammal carcasses are an important alternative without sea ice, especially carcasses of the
beluga whale The beluga whale () (''Delphinapterus leucas'') is an Arctic and sub-Arctic cetacean. It is one of two members of the family Monodontidae, along with the narwhal, and the only member of the genus ''Delphinapterus''. It is also known as the ...
. These alternatives may reduce the rate of weight loss of bears when on land. One scientist found that 71% of the
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
bears had fed on seaweed ( marine algae) and that about half were feeding on birds such as the dovekie and sea ducks, especially the long-tailed duck (53%) and common eider, by swimming underwater to catch them. They were also diving to feed on blue mussels and other underwater food sources like the green sea urchin. 24% had eaten moss recently, 19% had consumed grass, 34% had eaten black crowberry and about half had consumed willows. This study illustrates the polar bear's dietary flexibility but it does not represent its life history elsewhere. Most polar bears elsewhere will never have access to these alternatives, except for the marine mammal carcasses that are important wherever they occur. In Svalbard, polar bears were observed to kill white-beaked dolphins during spring, when the dolphins were trapped in the sea ice. The bears then proceeded to cache the carcasses, which remained and were eaten during the ice-free summer and autumn.


Reproduction and lifecycle

Courtship and mating take place on the sea ice in April and May, when polar bears congregate in the best seal hunting areas. A male may follow the tracks of a breeding female for or more, and after finding her engage in intense fighting with other males over mating rights, fights that often result in scars and broken teeth. Polar bears have a generally polygynous mating system; recent genetic testing of mothers and cubs, however, has uncovered cases of litters in which cubs have different fathers. Partners stay together and mate repeatedly for an entire week; the mating ritual induces ovulation in the female. Rosing, pp. 42–48 After mating, the fertilized egg remains in a suspended state until August or September. During these four months, the pregnant female eats prodigious amounts of food, gaining at least and often more than doubling her body weight.


Maternity denning and early life

When the ice floes are at their minimum in the fall, ending the possibility of hunting, each pregnant female digs a ''maternity den'' consisting of a narrow entrance tunnel leading to one to three chambers. Most maternity dens are in snowdrifts, but may also be made underground in permafrost if it is not sufficiently cold yet for snow. In most subpopulations, maternity dens are situated on land a few kilometres from the coast, and the individuals in a subpopulation tend to reuse the same denning areas each year. The polar bears that do not den on land make their dens on the sea ice. In the den, she enters a dormant state similar to
hibernation Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It mos ...
. This hibernation-like state does not consist of continuous sleeping; however, the bear's heart rate slows from 46 to 27 beats per minute. Lockwood, pp.17–21 Her body temperature does not decrease during this period as it would for a typical mammal in hibernation. Between November and February, cubs are born blind, covered with a light down fur, and weighing less than , but in captivity they might be delivered in the earlier months. The earliest recorded birth of polar bears in captivity was on 11 October 2011 in the Toronto Zoo. On average, each litter has two cubs. The family remains in the den until mid-February to mid-April, with the mother maintaining her fast while nursing her cubs on a fat-rich milk. By the time the mother breaks open the entrance to the den, her cubs weigh about . For about 12 to 15 days, the family spends time outside the den while remaining in its vicinity, the mother grazing on vegetation while the cubs become used to walking and playing. Then they begin the long walk from the denning area to the sea ice, where the mother can once again catch seals. Depending on the timing of ice-floe breakup in the fall, she may have fasted for up to eight months. During this time, cubs playfully imitate the mother's hunting methods in preparation for later life. Female polar bears have been known to adopt other cubs. Multiple cases of adoption of wild cubs have been confirmed by genetic testing. Adult bears of either sex occasionally kill and eat polar bear cubs. As of 2006, in Alaska, 42% of cubs were reaching 12 months of age, down from 65% in 1991. In most areas, cubs are weaned at two and a half years of age, when the mother chases them away or abandons them. The Western Hudson Bay subpopulation is unusual in that its female polar bears sometimes wean their cubs at only one and a half years. This was the case for 40% of cubs there in the early 1980s; however by the 1990s, fewer than 20% of cubs were weaned this young. After the mother leaves, sibling cubs sometimes travel and share food together for weeks or months.


Later life

Females begin to breed at the age of four years in most areas, and five years in the area of the Beaufort Sea. Males usually reach sexual maturity at six years; however, as competition for females is fierce, many do not breed until the age of eight or ten. A study in Hudson Bay indicated that both the reproductive success and the maternal weight of females peaked in their mid-teens. Maternal success appeared to decline after this point, possibly because of an age-related impairment in the ability to store the fat necessary to rear cubs. Polar bears appear to be less affected by infectious diseases and parasites than most terrestrial mammals. Polar bears are especially susceptible to '' Trichinella'', a parasitic roundworm they contract through cannibalism, although infections are usually not fatal. Only one case of a polar bear with
rabies Rabies is a viral disease that causes encephalitis in humans and other mammals. Early symptoms can include fever and tingling at the site of exposure. These symptoms are followed by one or more of the following symptoms: nausea, vomiting, ...
has been documented, even though polar bears frequently interact with Arctic foxes, which often carry rabies. Bacterial leptospirosis and '' Morbillivirus'' have been recorded. Polar bears sometimes have problems with various skin diseases that may be caused by
mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear e ...
s or other parasites.


Life expectancy

Polar bears rarely live beyond 25 years. Hemstock, pp. 29–35 The oldest wild bears on record died at age 32, whereas the oldest captive was a female who died in 1991, age 43. The causes of death in wild adult polar bears are poorly understood, as carcasses are rarely found in the species's frigid habitat. In the wild, old polar bears eventually become too weak to catch food, and gradually starve to death. Polar bears injured in fights or accidents may either die from their injuries, or become unable to hunt effectively, leading to starvation.


Ecological role

The polar bear is the apex predator within its range, and is a keystone species for the Arctic. Several animal species, particularly Arctic foxes (''Vulpes lagopus'') and glaucous gulls (''Larus hyperboreus''), routinely scavenge polar bear kills. The relationship between ringed seals and polar bears is so close that the abundance of ringed seals in some areas appears to regulate the density of polar bears, while polar bear predation in turn regulates density and reproductive success of ringed seals. The evolutionary pressure of polar bear predation on seals probably accounts for some significant differences between Arctic and Antarctic seals. Compared to the
Antarctic The Antarctic ( or , American English also or ; commonly ) is a polar region around Earth's South Pole, opposite the Arctic region around the North Pole. The Antarctic comprises the continent of Antarctica, the Kerguelen Plateau and othe ...
, where there is no major surface predator, Arctic seals use more breathing holes per individual, appear more restless when hauled out on the ice, and rarely defecate on the ice. The baby fur of most Arctic seal species is white, presumably to provide camouflage from predators, whereas Antarctic seals all have dark fur at birth. Brown bears tend to dominate polar bears in disputes over carcasses, and dead polar bear cubs have been found in brown bear dens. Wolves are rarely encountered by polar bears, though there are two records of Arctic wolf (''Canis lupus arctos'') packs killing polar bear cubs. Adult polar bears are occasionally vulnerable to predation by
orca The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus '' Orcinus'' and is recognizable by its black-and-white ...
s (''Orcinus orca'') while swimming, but they are rarely reported as taken and bears are likely to avoid entering the water if possible if they detect an orca pod in the area. The melting sea ice in the Arctic may be causing an increase of orcas in the Arctic sea, which may increase the risk of predation on polar bears but also may benefit the bears by providing more whale carcasses that they can scavenge. The remains of polar bears have been found in the stomachs of large
Greenland shark The Greenland shark (''Somniosus microcephalus''), also known as the gurry shark, grey shark, or by the Kalaallisut name ''eqalussuaq'', is a large shark of the family Somniosidae ("sleeper sharks"), closely related to the Pacific and souther ...
s (''Somniosus microcephalus''), although it certainly cannot be ruled out that the bears were merely scavenged by this slow-moving, unusual shark. A rather unlikely killer of a grown polar bear has reportedly included a
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for " glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is a musc ...
(''Gulo gulo''), anecdotally reported to have suffocated a bear in a zoo with a bite to the throat during a conflict. This report may well be dubious, however. Polar bears are sometimes the host of arctic mites such as ''
Alaskozetes antarcticus ''Alaskozetes antarcticus'' is a species of non-parasitic mite, known for its ability to survive in subzero temperatures. This animal's name derives from its habitat: ''Alasko'', meaning "from Alaska", and ''antarcticus'', in connection with fr ...
''.


Long-distance swimming and diving

Researchers tracked 52 sows in the southern Beaufort Sea off Alaska with GPS system collars; no boars were involved in the study due to males' necks being too thick for the GPS-equipped collars. Fifty long-distance swims were recorded; the longest at , with an average of . The length of these swims ranged from most of a day to ten days. Ten of the sows had a cub swim with them and after a year, six cubs survived. The study did not determine if the others lost their cubs before, during, or some time after their long swims. Researchers do not know whether or not this is a new behaviour; before polar ice shrinkage, they opined that there was probably neither the need nor opportunity to swim such long distances. As noted above, there have been 600 recorded instances of polar bears in Iceland since the 9th century. These presumably swam from Greenland, about . The polar bear may swim underwater for up to three minutes to approach seals on shore or on ice floes.


Hunting


Indigenous people

Polar bears have long provided important raw materials for Arctic peoples, including the Inuit, Yupik, Chukchi, Nenets, Russian Pomors and others. Hunters commonly used teams of dogs to distract the bear, allowing the hunter to spear the bear or shoot it with arrows at closer range. Almost all parts of captured animals had a use. The fur was used in particular to make trousers and, by the Nenets, to make galoshes-like outer footwear called ; the meat is edible, despite some risk of trichinosis; the fat was used in food and as a fuel for lighting homes, alongside seal and whale blubber;
sinews A tendon or sinew is a tough, high-tensile-strength band of dense fibrous connective tissue that connects muscle to bone. It is able to transmit the mechanical forces of muscle contraction to the skeletal system without sacrificing its abilit ...
were used as thread for sewing clothes; the gallbladder and sometimes
heart The heart is a muscular Organ (biology), organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as ca ...
were dried and powdered for medicinal purposes; the large canine teeth were highly valued as
talismans A talisman is any object ascribed with religious or magical powers intended to protect, heal, or harm individuals for whom they are made. Talismans are often portable objects carried on someone in a variety of ways, but can also be installed perm ...
. Only the
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
was not used, as its high concentration of vitamin A is poisonous. As a carnivore, which feeds largely upon
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
-eating carnivores, the polar bear ingests large amounts of vitamin A that is stored in their
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
s. The resulting high concentrations cause
hypervitaminosis A Hypervitaminosis A refers to the toxic effects of ingesting too much preformed vitamin A (retinyl esters, retinol, and retinal). Symptoms arise as a result of altered bone metabolism and altered metabolism of other fat-soluble vitamins. Hypervi ...
, Hunters make sure to either toss the liver into the sea or bury it in order to spare their dogs from potential poisoning. Traditional subsistence hunting was on a small enough scale to not significantly affect polar bear populations, mostly because of the sparseness of the human population in polar bear habitat.


History of commercial harvest

In Russia, polar bear furs were already being commercially traded in the 14th century, though it was of low value compared to Arctic fox or even reindeer fur. The growth of the human population in the Eurasian Arctic in the 16th and 17th century, together with the advent of firearms and increasing trade, dramatically increased the harvest of polar bears. However, since polar bear fur has always played a marginal commercial role, data on the historical harvest is fragmentary. It is known, for example, that already in the winter of 1784/1785 Russian Pomors on Spitsbergen harvested 150 polar bears in Magdalenefjorden. In the early 20th century, Norwegian hunters were harvesting 300 bears per year at the same location. Estimates of total historical harvest suggest that from the beginning of the 18th century, roughly 400 to 500 animals were being harvested annually in northern Eurasia, reaching a peak of 1,300 to 1,500 animals in the early 20th century, and falling off as the numbers began dwindling. In the first half of the 20th century, mechanized and overpoweringly efficient methods of hunting and trapping came into use in North America as well. Polar bears were chased from snowmobiles, icebreakers, and airplanes, the latter practice described in a 1965 ''New York Times'' editorial as being "about as sporting as machine gunning a cow." Bruemmer, pp. 93–111 Norwegians used "self-killing guns", comprising a loaded rifle in a baited box that was placed at the level of a bear's head, and which fired when the string attached to the bait was pulled. The numbers taken grew rapidly in the 1960s, peaking around 1968 with a global total of 1,250 bears that year.


Contemporary regulations

Concerns over the future survival of the species led to the development of national regulations on polar bear hunting, beginning in the mid-1950s. The Soviet Union banned all hunting in 1956. Canada began imposing hunting quotas in 1968. Norway passed a series of increasingly strict regulations from 1965 to 1973, and has completely banned hunting since then. The United States began regulating hunting in 1971 and adopted the Marine Mammal Protection Act in 1972. In 1973, the
International Agreement on the Conservation of Polar Bears The Agreement on the Conservation of Polar Bears is a multilateral treaty signed in Oslo, November 15, 1973, by the five nations with the largest polar bear populations: Canada, Denmark ( Greenland), Norway ( Svalbard), the United States, and the ...
was signed by all five nations whose territory is inhabited by polar bears: Canada, Denmark, Norway, the
Soviet Union The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, it was nominally a federal union of fifteen nationa ...
, and the United States. Member countries agreed to place restrictions on recreational and commercial hunting, ban hunting from aircraft and icebreakers, and conduct further research. The treaty allows hunting "by local people using traditional methods". Norway is the only country of the five in which all harvest of polar bears is banned. The agreement was a rare case of international cooperation during the
Cold War The Cold War is a term commonly used to refer to a period of geopolitical tension between the United States and the Soviet Union and their respective allies, the Western Bloc and the Eastern Bloc. The term '' cold war'' is used because t ...
. Biologist Ian Stirling commented, "For many years, the conservation of polar bears was the only subject in the entire Arctic that nations from both sides of the Iron Curtain could agree upon sufficiently to sign an agreement. Such was the intensity of human fascination with this magnificent predator, the only marine bear." Agreements have been made between countries to co-manage their shared polar bear subpopulations. After several years of negotiations, Russia and the United States signed an agreement in October 2000 to jointly set quotas for indigenous subsistence hunting in Alaska and Chukotka. The treaty was ratified in October 2007. In September 2015, the polar bear range states agreed upon a "circumpolar action plan" describing their conservation strategy for polar bears. The species is listed in Appendix II of the
Convention on International Trade in Endangered Species CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
(CITES) meaning international trade, including in parts or derivatives, is controlled by the CITES system of permits and certificates. The United States government has proposed that polar bears be transferred to Appendix I of
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
which would ban all commercial international trade in polar bear parts. The decision to leave the species listed under Appendix II was endorsed by the IUCN and
TRAFFIC Traffic comprises pedestrians, vehicles, ridden or herded animals, trains, and other conveyances that use public ways (roads) for travel and transportation. Traffic laws govern and regulate traffic, while rules of the road include traffic ...
, who determined that such an uplisting was unlikely to confer a conservation benefit.


Canada

Polar bears were designated "Not at Risk" in April 1986 and uplisted to "Special Concern" in April 1991. This status was re-evaluated and confirmed in April 1999, November 2002, and April 2008. Polar bears continue to be listed as a species of special concern in Canada because of their sensitivity to overharvest and because of an expected range contraction caused by loss of Arctic sea ice. More than 600 bears are killed per year by humans across Canada, a rate calculated by scientists to be unsustainable for some areas, notably Baffin Bay. Canada has allowed sport hunters accompanied by local guides and dog-sled teams since 1970, but the practice was not common until the 1980s. The guiding of sport hunters provides meaningful employment and an important source of income for northern communities in which economic opportunities are few. Sport hunting can bring CDN$20,000 to $35,000 per bear into northern communities, which until recently has been mostly from American hunters. The territory of
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
accounts for the location 80% of annual kills in Canada. In 2005, the government of Nunavut increased the quota from 400 to 518 bears, despite protests from the IUCN Polar Bear Specialist Group. In two areas where harvest levels have been increased based on increased sightings, science-based studies have indicated declining populations, and a third area is considered data-deficient. While most of that quota is hunted by the indigenous Inuit, a growing share is sold to recreational hunters. (0.8% in the 1970s, 7.1% in the 1980s, and 14.6% in the 1990s) Nunavut polar bear biologist, Mitchell Taylor, who was formerly responsible for polar bear conservation in the territory, has insisted that bear numbers are being sustained under current hunting limits. In 2010, the 2005 increase was partially reversed. Government of Nunavut officials announced that the polar bear quota for the Baffin Bay region would be gradually reduced from 105 per year to 65 by 2013. The Government of the
Northwest Territories The Northwest Territories (abbreviated ''NT'' or ''NWT''; french: Territoires du Nord-Ouest, formerly ''North-Western Territory'' and ''North-West Territories'' and namely shortened as ''Northwest Territory'') is a federal territory of Canada. ...
maintain their own quota of 72 to 103 bears within the Inuvialuit communities of which some are set aside for sports hunters.
Environment Canada Environment and Climate Change Canada (ECCC; french: Environnement et Changement climatique Canada),Environment and Climate Change Canada is the applied title under the Federal Identity Program; the legal title is Department of the Environment ...
also banned the export from Canada of fur, claws, skulls and other products from polar bears harvested in Baffin Bay as of 1 January 2010. Because of the way polar bear hunting quotas are managed in Canada, attempts to discourage sport hunting would actually increase the number of bears killed in the short term. Canada allocates a certain number of permits each year to sport and subsistence hunting, and those that are not used for sport hunting are re-allocated to indigenous subsistence hunting. Whereas northern communities kill all the polar bears they are permitted to take each year, only half of sport hunters with permits actually manage to kill a polar bear. If a sport hunter does not kill a polar bear before his or her permit expires, the permit cannot be transferred to another hunter. In August 2011,
Environment Canada Environment and Climate Change Canada (ECCC; french: Environnement et Changement climatique Canada),Environment and Climate Change Canada is the applied title under the Federal Identity Program; the legal title is Department of the Environment ...
published a national polar bear conservation strategy.


Greenland

In Greenland, hunting restrictions were first introduced in 1994 and expanded by executive order in 2005. Until 2005 Greenland placed no limit on hunting by indigenous people. However, in 2006 it imposed a limit of 150, while also allowed recreational hunting for the first time. Other provisions included year-round protection of cubs and mothers, restrictions on weapons used and various administrative requirements to catalogue kills.


Norway

Polar bears were hunted heavily in Svalbard, Norway throughout the 19th century and to as recently as 1973, when the conservation treaty was signed. 900 bears a year were harvested in the 1920s and after World War II, there were as many as 400–500 harvested annually. Some regulations of hunting did exist. In 1927, poisoning was outlawed while in 1939, certain denning sights were declared off limits. The killing of females and cubs was made illegal in 1965. Killing of polar bears decreased somewhat 25–30 years before the treaty. Despite this, the polar bear population continued to decline and by 1973, only around 1000 bears were left in Svalbard. Only with the passage of the treaty did they begin to recover.


Russia

The Soviet Union banned the harvest of polar bears in 1956; however, poaching continued, and is estimated to pose a serious threat to the polar bear population. In recent years, polar bears have approached coastal villages in Chukotka more frequently due to the shrinking of the sea ice, endangering humans and raising concerns that illegal hunting would become even more prevalent. In 2007, the Russian government made subsistence hunting legal for indigenous Chukotkan peoples only, a move supported by Russia's most prominent bear researchers and the
World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the W ...
as a means to curb poaching. Polar bears are currently listed as "Rare", of "Uncertain Status", or "Rehabilitated and rehabilitating" in the Red Data Book of Russia, depending on population. In 2010, the Ministry of Natural Resources and Environment published a strategy for polar bear conservation in Russia.


United States

The Marine Mammal Protection Act of 1972 afforded polar bears some protection in the United States. It banned hunting (except by indigenous subsistence hunters), banned importing of polar bear parts (except polar bear pelts taken legally in Canada), and banned the harassment of polar bears. On 15 May 2008, the
United States Department of the Interior The United States Department of the Interior (DOI) is one of the executive departments of the U.S. federal government headquartered at the Main Interior Building, located at 1849 C Street NW in Washington, D.C. It is responsible for the ma ...
listed the polar bear as a threatened species under the Endangered Species Act, citing the melting of Arctic sea ice as the primary threat to the polar bear. It banned all importing of polar bear trophies. Importing products made from polar bears had been prohibited from 1972 to 1994 under the Marine Mammal Protection Act, and restricted between 1994 and 2008. Under those restrictions, permits from the United States Fish and Wildlife Service were required to import sport-hunted polar bear trophies taken in hunting expeditions in Canada. The permit process required that the bear be taken from an area with quotas based on sound management principles. Since 1994, hundreds of sport-hunted polar bear trophies have been imported into the U.S. In 2015, the U.S. Fish and Wildlife Service published a draft conservation management plan for polar bears to improve their status under the Endangered Species Act and the Marine Mammal Protection Act.


Conservation status, threats, and controversies

Polar bear population sizes and trends are difficult to estimate accurately because they occupy remote home ranges and exist at low population densities. Polar bear fieldwork can also be hazardous to researchers. As of 2015, the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) reports that the global population of polar bears is 22,000 to 31,000, and the current population trend is unknown. Nevertheless, polar bears are listed as "Vulnerable" under criterion A3c, which indicates an expected population decrease of ≥30% over the next three generations (~34.5 years) due to "decline in area of occupancy, extent of occurrence and/or quality of habitat". Risks to the polar bear include
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
,
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, th ...
in the form of
toxic Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a sub ...
contaminants, conflicts with shipping, oil and gas exploration and development, and human-bear interactions including harvesting and possible stresses from recreational polar-bear watching. According to the
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the W ...
, the polar bear is important as an
indicator Indicator may refer to: Biology * Environmental indicator of environmental health (pressures, conditions and responses) * Ecological indicator of ecosystem health (ecological processes) * Health indicator, which is used to describe the health o ...
of Arctic ecosystem health. Polar bears are studied to gain understanding of what is happening throughout the Arctic, because at-risk polar bears are often a sign of something wrong with the Arctic marine ecosystem.


Climate change

The key danger for polar bears posed by the
effects of climate change The effects of climate change impact the physical environment, ecosystems and human societies. The environmental effects of climate change are broad and far-reaching. They affect the water cycle, oceans, sea and land ice ( glaciers), sea le ...
is malnutrition or starvation due to
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
. Polar bears hunt seals from a platform of sea ice. Rising temperatures cause the sea ice to melt earlier in the year, driving the bears to shore before they have built sufficient fat reserves to survive the period of scarce food in the late summer and early fall. Reduction in sea-ice cover also forces bears to swim longer distances, which further depletes their energy stores and occasionally leads to drowning. Thinner sea ice tends to deform more easily, which appears to make it more difficult for polar bears to access seals. Insufficient nourishment leads to lower reproductive rates in adult females and lower survival rates in cubs and juvenile bears, in addition to poorer body condition in bears of all ages. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, Arctic Climate Impact Assessment,
United States Geological Survey The United States Geological Survey (USGS), formerly simply known as the Geological Survey, is a scientific agency of the United States government. The scientists of the USGS study the landscape of the United States, its natural resources, ...
and many leading polar bear biologists have expressed grave concerns about the impact of climate change, with some predicting extinction by 2100. In addition to creating nutritional stress, a warming climate is expected to affect various other aspects of polar bear life: changes in sea ice affect the ability of pregnant females to build suitable maternity dens. As the distance increases between the pack ice and the coast, females must swim longer distances to reach favoured denning areas on land. Thawing of permafrost would affect the bears who traditionally den underground, and warm winters could result in den roofs collapsing or having reduced insulative value. For the polar bears that currently den on multi-year ice, increased ice mobility may result in longer distances for mothers and young cubs to walk when they return to seal-hunting areas in the spring. Disease-causing
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
and
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson h ...
s would flourish more readily in a warmer climate. Problematic interactions between polar bears and humans, such as foraging by bears in garbage dumps, have historically been more prevalent in years when ice-floe breakup occurred early and local polar bears were relatively thin. Increased human-bear interactions, including fatal attacks on humans, are likely to increase as the sea ice shrinks and hungry bears try to find food on land. The
effects of climate change The effects of climate change impact the physical environment, ecosystems and human societies. The environmental effects of climate change are broad and far-reaching. They affect the water cycle, oceans, sea and land ice ( glaciers), sea le ...
are most profound in the southern part of the polar bear's range, and this is indeed where significant degradation of local populations has been observed.. The Western Hudson Bay subpopulation, in a southern part of the range, also happens to be one of the best-studied polar bear subpopulations. This subpopulation feeds heavily on ringed seals in late spring, when newly weaned and easily hunted seal pups are abundant. The late spring hunting season ends for polar bears when the ice begins to melt and break up, and they fast or eat little during the summer until the sea freezes again. Due to warming air temperatures, ice-floe breakup in western Hudson Bay is currently occurring three weeks earlier than it did 30 years ago, reducing the duration of the polar bear feeding season. The body condition of polar bears has declined during this period; the average weight of lone (and likely pregnant) female polar bears was approximately in 1980 and in 2004. Between 1987 and 2004, the Western Hudson Bay population declined by 22%, although the population was listed as "stable" as of 2017. As the climate change melts sea ice, the U.S. Geological Survey projects that two-thirds of polar bears will disappear by 2050. In Alaska, the effects of sea ice shrinkage have contributed to higher mortality rates in polar bear cubs, and have led to changes in the denning locations of pregnant females. The proportion of maternity dens on sea ice has changed from 62% between the years 1985 through 1994, to 37% over the years 1998 through 2004. Thus, now the Alaskan population more resembles the world population in that it is more likely to den on land. In recent years, polar bears in the Arctic have undertaken longer than usual swims to find prey, possibly resulting in four recorded drownings in the unusually large ice pack regression of 2005. A new development is that polar bears have begun ranging to new territory. While not unheard of but still uncommon, polar bears have been sighted increasingly in larger numbers ashore, staying on the mainland for longer periods of time during the summer months, particularly in North Canada, traveling farther inland. This may cause an increased reliance on terrestrial diets, such as goose eggs, waterfowl and caribou, as well as increased human–bear conflict.


Pollution

Polar bears accumulate high levels of persistent organic pollutants such as
polychlorinated biphenyl Polychlorinated biphenyls (PCBs) are highly carcinogenic chemical compounds, formerly used in industrial and consumer products, whose production was banned in the United States by the Toxic Substances Control Act of 1976, Toxic Substances Contro ...
(PCBs) and chlorinated pesticides. Due to their position at the top of the
ecological pyramid An ecological pyramid (also trophic pyramid, Eltonian pyramid, energy pyramid, or sometimes food pyramid) is a graphical representation designed to show the biomass or bioproductivity at each trophic level in a given ecosystem. A ''pyramid of ...
, with a diet heavy in blubber in which halocarbons concentrate, their bodies are among the most contaminated of Arctic mammals. Halocarbons (also known as organohalogens) are known to be toxic to other animals, because they mimic
hormone A hormone (from the Greek participle , "setting in motion") is a class of signaling molecules in multicellular organisms that are sent to distant organs by complex biological processes to regulate physiology and behavior. Hormones are required ...
chemistry, and biomarkers such as immunoglobulin G and retinol suggest similar effects on polar bears. PCBs have received the most study, and they have been associated with birth defects and immune system deficiency. Many chemicals, such as PCBs and DDT, have been internationally banned due to the recognition of their harm on the environment. Their concentrations in polar bear tissues continued to rise for decades after being banned, as these chemicals spread through the food chain. Since then, the trend seems to have abated, with tissue concentrations of PCBs declining between studies performed from 1989 to 1993 and studies performed from 1996 to 2002. During the same time periods, DDT was found to be notably lower in the Western Hudson Bay population only.


Oil and gas development

Oil and gas development in polar bear habitat can affect the bears in a variety of ways. An oil spill in the Arctic would most likely concentrate in the areas where polar bears and their prey are also concentrated, such as sea ice leads. Because polar bears rely partly on their fur for insulation and soiling of the fur by oil reduces its insulative value, oil spills put bears at risk of dying from
hypothermia Hypothermia is defined as a body core temperature below in humans. Symptoms depend on the temperature. In mild hypothermia, there is shivering and mental confusion. In moderate hypothermia, shivering stops and confusion increases. In severe ...
. Polar bears exposed to oil spill conditions have been observed to lick the oil from their fur, leading to fatal kidney failure. Maternity dens, used by pregnant females and by females with infants, can also be disturbed by nearby oil exploration and development. Disturbance of these sensitive sites may trigger the mother to abandon her den prematurely, or abandon her litter altogether.


Predictions

Steven Amstrup Steven C. Amstrup (born February 4, 1950) is an American zoologist who studies bears, especially polar bears. He is the 2012 recipient of the Indianapolis Prize. Early life Steven Amstrup was born in Fargo, North Dakota, where he took an intere ...
and other U.S. Geological Survey scientists have predicted two-thirds of the world's polar bears may disappear by 2050, based on moderate projections for the shrinking of summer sea ice caused by climate change, though the validity of this study has been debated. The bears could disappear from Europe, Asia, and Alaska, and be depleted from the Canadian Arctic Archipelago and areas off the northern Greenland coast. By 2080, they could disappear from Greenland entirely and from the northern Canadian coast, leaving only dwindling numbers in the interior Arctic Archipelago. However, in the short term, some polar bear populations in historically colder regions of the Arctic may temporarily benefit from a milder climate, as multiyear ice that is too thick for seals to create breathing holes is replaced by thinner annual ice. Polar bears diverged from brown bears 400,000–600,000 years ago and have survived past periods of climate fluctuation. It has been claimed that polar bears will be able to adapt to terrestrial food sources as the sea ice they use to hunt seals disappears. However, most polar bear biologists think that polar bears will be unable to completely offset the loss of calorie-rich seal blubber with terrestrial foods, and that they will be outcompeted by brown bears in this terrestrial niche, ultimately leading to a population decline.


Controversy over species protection

Warnings about the future of the polar bear are often contrasted with the fact that worldwide population estimates have increased over the past 50 years and are relatively stable today. Some estimates of the global population are around 5,000 to 10,000 in the early 1970s; other estimates were 20,000 to 40,000 during the 1980s. Current estimates put the global population at between 20,000 and 25,000 or 22,000 and 31,000. Despite the encouraging rebound of some populations, there is little evidence to suggest polar bears are thriving overall. There are several reasons for the apparent discordance between past and projected population trends: estimates from the 1950s and 1960s were based on stories from explorers and hunters rather than on scientific surveys. Second, controls of harvesting were introduced that allowed this previously overhunted species to recover. Third, the recent effects of climate change have affected sea ice abundance in different areas to varying degrees. Debate over the listing of the polar bear under endangered species legislation has put conservation groups and Canada's
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...
at opposing positions; the
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
government and many northern residents have condemned the U.S. initiative to list the polar bear under the Endangered Species Act. Many Inuit believe the polar bear population is increasing, and restrictions on commercial sport-hunting are likely to lead to a loss of income to their communities.Northern Research Forum.
Polar bear as a resource
'' A position paper presented for the 3rd NRF Open Meeting in Yellowknife and Rae Edzo, Canada. 15–18 September 2004


In culture


Indigenous folklore

For the indigenous peoples of the Arctic, polar bears have long played an important cultural and material role. Lockwood, pp. 6–9 Polar bear remains have been found at hunting sites dating to 2,500 to 3,000 years ago Lockwood, pp. 31–36 and 1,500-year-old cave paintings of polar bears have been found in the Chukchi Peninsula. Indeed, it has been suggested that Arctic peoples' skills in seal hunting and igloo construction has been in part acquired from the polar bears themselves. The
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...
and Alaska Natives have many folk tales featuring the bears including legends in which bears are humans when inside their own houses and put on bear hides when going outside, and stories of how the constellation that is said to resemble a great bear surrounded by dogs came into being. These legends reveal a deep respect for the polar bear, which is portrayed as both spiritually powerful and closely akin to humans. The human-like posture of bears when standing and sitting, and the resemblance of a skinned bear carcass to the human body, have probably contributed to the belief that the spirits of humans and bears were interchangeable. Among the Chukchi and Yupik of eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
, there was a longstanding shamanistic ritual of "thanksgiving" to the hunted polar bear. After killing the animal, its head and skin were removed and cleaned and brought into the home, and a feast was held in the hunting camp in its honor. To appease the spirit of the bear, traditional song and drum music was played, and the skull was ceremonially fed and offered a pipe. Only once the spirit was appeased was the skull be separated from the skin, taken beyond the bounds of the homestead, and placed in the ground, facing north. The Nenets of north-central Siberia placed particular value on the talismanic power of the prominent canine teeth. These were traded in the villages of the lower Yenisei and Khatanga rivers to the forest-dwelling peoples further south, who would sew them into their hats as protection against
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
s. It was believed that the "little nephew" (the brown bear) would not dare to attack a man wearing the tooth of its powerful "big uncle", the polar bear. The skulls of killed polar bears were buried at sacred sites, and altars, called ''sedyangi'', were constructed out of the skulls. Several such sites have been preserved on the Yamal Peninsula.


Symbols and mascots

Their distinctive appearance and their association with the Arctic have made polar bears popular icons, especially in those areas where they are native. The Canadian two-dollar coin carries an image of a lone polar bear on its reverse side, while a special millennium edition featured three. Vehicle licence plates in the
Northwest Territories The Northwest Territories (abbreviated ''NT'' or ''NWT''; french: Territoires du Nord-Ouest, formerly ''North-Western Territory'' and ''North-West Territories'' and namely shortened as ''Northwest Territory'') is a federal territory of Canada. ...
in Canada are in the shape of a polar bear, as was the case in
Nunavut Nunavut ( , ; iu, ᓄᓇᕗᑦ , ; ) is the largest and northernmost territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the '' Nunavut Act'' and the '' Nunavut Land Claims Agreement Act'' ...
until 2012; these now display polar bear artwork instead. The polar bear is the mascot of
Bowdoin College Bowdoin College ( ) is a private liberal arts college in Brunswick, Maine. When Bowdoin was chartered in 1794, Maine was still a part of the Commonwealth of Massachusetts. The college offers 34 majors and 36 minors, as well as several joint eng ...
,
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
; the
University of Alaska Fairbanks The University of Alaska Fairbanks (UAF or Alaska) is a public land-grant research university in College, Alaska, a suburb of Fairbanks. It is the flagship campus of the University of Alaska system. UAF was established in 1917 and opened for c ...
; and the 1988 Winter Olympics held in
Calgary Calgary ( ) is the largest city in the western Canadian province of Alberta and the largest metro area of the three Prairie Provinces. As of 2021, the city proper had a population of 1,306,784 and a metropolitan population of 1,481,806, maki ...
. The Eisbären Berlin hockey team uses a roaring polar bear as their logo, and the
Charlotte, North Carolina Charlotte ( ) is the most populous city in the U.S. state of North Carolina. Located in the Piedmont region, it is the county seat of Mecklenburg County. The population was 874,579 at the 2020 census, making Charlotte the 16th-most popu ...
hockey team the
Charlotte Checkers The Charlotte Checkers are a minor-league professional ice hockey team based in Charlotte, North Carolina. They are members of the Atlantic Division of the Eastern Conference in the American Hockey League (AHL), and are the top minor league aff ...
uses a polar bear named Chubby Checker as their mascot.
Coca-Cola Coca-Cola, or Coke, is a carbonated soft drink manufactured by the Coca-Cola Company. Originally marketed as a temperance bar, temperance drink and intended as a patent medicine, it was invented in the late 19th century by John Stith Pembe ...
has used images of the polar bear in its advertising, and Polar Beverages, Nelvana,
Bundaberg Rum Bundaberg Rum, colloquially known as Bundy, is a dark rum produced in Bundaberg East, Queensland, Australia, by the Bundaberg Distilling Company. In 2010, the Bundaberg Distilling Company was inducted into the Queensland Business Leaders Hall ...
, Klondike bars, and
Fox's Glacier Mints Fox's Glacier Mints are a brand of boiled mint sold in the UK since 1918. History Background Walter Richard Fox, the founder of Fox's Confectionery, originally manufactured sweets as part of the Joyce and Fox partnership, but this was dissolved ...
all feature polar bears in their logos.


Fiction

Polar bears are popular in fiction, particularly in books for children or teenagers. *''The Polar Bear Son'' is adapted from a traditional Inuit tale. *The animated television series '' Noah's Island'' features a polar bear named Noah as the protagonist. *Polar bears feature prominently in '' East'' (''North Child'' in the UK) by Edith Pattou, *''The Bear'' by Raymond Briggs (adapted into an animated short in 1998) is a story of a girl's encounter with a polar bear. *The '' panserbjørne'' of Philip Pullman's fantasy trilogy '' His Dark Materials'' are sapient, dignified,
anthropomorphic Anthropomorphism is the attribution of human traits, emotions, or intentions to non-human entities. It is considered to be an innate tendency of human psychology. Personification is the related attribution of human form and characteristics t ...
polar bears. They feature prominently in the 2007 film adaptation of ''The Golden Compass''. *The television series ''
Lost Lost may refer to getting lost, or to: Geography * Lost, Aberdeenshire, a hamlet in Scotland *Lake Okeechobee Scenic Trail, or LOST, a hiking and cycling trail in Florida, US History *Abbreviation of lost work, any work which is known to have bee ...
'' features polar bears living on the tropical island setting. *The manga and anime series ''
Shirokuma Cafe is a Japanese manga series by Aloha Higa. It revolves around the everyday lives of a group of animals mingling with humans at a café run by a polar bear. An anime adaptation by Studio Pierrot aired in Japan between April 2012 and March 201 ...
'' centers on a polar bear who runs a cafe in Japan.


See also

* Charismatic megafauna * Flagship species *
International Polar Bear Day International Polar Bear Day is an annual event celebrated every February 27, to coincide with the time period when polar bear moms and cubs are sleeping in their dens, and to raise awareness about the conservation status of the polar bear. Descr ...
*
2019 mass invasion of Russian polar bears In February 2019, the Russian archipelago of Novaya Zemlya in the Arctic Ocean experienced a mass invasion of polar bears. Dozens of polar bears were seen trying to enter homes, civic buildings, and inhabited areas. The Arkhangelsk Oblast author ...
* Umbrella species


References


Bibliography

* * * * * *


Further reading

* .


External links


National Wildlife Federation's Polar Bear Page
* ARKive 
images and movies of the polar bear (''Ursus maritimus'')

Map of polar bear ranges and denning areas in Nunavut from Nunavut Planning Commission
* BBC Nature
Polar bear news, and video clips from BBC programmes past and present.

Photos, facts, videos from Polar Bears International that funds population, preservation, and DNA studies of the polar bear

Map: Here's where the polar bears are vanishing
* {{DEFAULTSORT:Polar Bear Apex predators Pleistocene bears Articles containing video clips Carnivorans of Europe Carnivorans of North America Carnivorans of Asia Holarctic fauna Mammals of the Arctic Marine mammals Mammals described in 1774 Vulnerable animals Vulnerable biota of Europe Vulnerable biota of Asia ESA threatened species Taxa named by Constantine Phipps, 2nd Baron Mulgrave Species endangered by climate change