Pied-billed grebe
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The pied-billed grebe (''Podilymbus podiceps'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the
grebe Grebes () are aquatic diving birds in the order Podicipediformes . Grebes are widely distributed freshwater birds, with some species also found in marine habitats during migration and winter. Some flightless species exist as well, most notably ...
family of water birds. Because the
Atitlán grebe The Atitlán grebe (''Podilymbus gigas''), also known as giant grebe, giant pied-billed grebe, or poc, is an extinct water bird, a relative of the pied-billed grebe. It was endemic at the Lago de Atitlán in Guatemala at an altitude of 1700 m ...
(''Podilymbus gigas'') has become
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
, the Pied-Billed Grebe is now the sole extant member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Podilymbus ''Podilymbus'' is a genus of birds in the Grebe family, the genus name is derived from Latin ''Podilymbus'', a contraction of ''podicipes'' ("feet at the buttocks", from ''podici-'', "rump-" + ''pes'', "foot")—the origin of the name of the g ...
''. The pied-billed grebe is primarily found in ponds throughout the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
. Other names of this grebe include American dabchick, rail, dabchick, Carolina grebe, devil-diver, dive-dapper, dipper, hell-diver, pied-billed dabchick, pied-bill, thick-billed grebe, and water witch.


Taxonomy and name

The pied-billed grebe was described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in the 10th edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' in 1758 as ''Colymbus podiceps''. The binomial name is derived from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''Podilymbus'', a
contraction Contraction may refer to: Linguistics * Contraction (grammar), a shortened word * Poetic contraction, omission of letters for poetic reasons * Elision, omission of sounds ** Syncope (phonology), omission of sounds in a word * Synalepha, merged ...
of ''podicipes'' ("feet at the buttocks", from ''podici-'', "rump-" + ''pes'', "foot")—the origin of the name of the grebe order—and
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''kolymbos'', "diver", and ''podiceps'', "rump-headed", from ''podici-'' +
New Latin New Latin (also called Neo-Latin or Modern Latin) is the revival of Literary Latin used in original, scholarly, and scientific works since about 1500. Modern scholarly and technical nomenclature, such as in zoological and botanical taxonomy ...
''ceps''. Outside its own genus, the closest relatives of the pied-billed grebe are the small grebes of the genus ''
Tachybaptus ''Tachybaptus'' is a genus of small members of the grebe family birds. The genus name means "quick diving": it is from Ancient Greek ''takhys'' "quick" and ''bapto'' "I dip". It has representatives over much of the world, including the tropics. ...
''.


Subspecies

*''P. p. podiceps'', (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
)
,
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, from
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
to
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
. *''P. p. antillarum'', (
Bangs Bang or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Lorestan, a village in I ...
, 1913)
, Greater &
Lesser Antilles The Lesser Antilles ( es, link=no, Antillas Menores; french: link=no, Petites Antilles; pap, Antias Menor; nl, Kleine Antillen) are a group of islands in the Caribbean Sea. Most of them are part of a long, partially volcanic island arc betwe ...
. *''P. p. antarcticus'', (
Lesson A lesson or class is a structured period of time where learning is intended to occur. It involves one or more students (also called pupils or learners in some circumstances) being taught by a teacher or instructor. A lesson may be either one ...
, 1842)
,
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, from
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
to central
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
&
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
.


Description

Pied-billed grebes are small, stocky, and short-necked. They are in length, with a wingspan of and weigh .Muller, M. J., and R. W. Storer. Pied-billed Grebe (''Podilymbus podiceps''). In The Birds of North America, No. 410 (A. Poole and F. Gill, eds.). Philadelphia, PA: The Birds of North America, Inc., 1999. They are mainly brown, with a darker crown and back. Their brown color serves as
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
in the marshes they live in. They do not have white visible under their wings when flying, like other grebes. Their undertail is white and they have a short, blunt chicken-like bill that is a light grey color, which in summer is encircled by a broad black band (hence the name). In the summer, its throat is black. There is no
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
. Juveniles have black and white stripes and look more like winter adults. This grebe does not have
webbed feet The webbed foot is a specialized limb with interdigital membranes (webbings) that aids in aquatic locomotion, present in a variety of tetrapod vertebrates. This adaptation is primarily found in semiaquatic species, and has convergently evolved m ...
. Its toes have
lobe Lobe may refer to: People with the name * Lobe (surname) Science and healthcare * Lobe (anatomy) * Lobe, a large-scale structure of a radio galaxy * Glacial lobe, a lobe-shaped glacier * Lobation, a characteristic of the nucleus of certain biolo ...
s that come out of the side of each toe. These lobes allow for easy paddling. When flying, the feet appear behind the body due to the feet's placement in the far back of the body. Because of the feet placement, they are not able to walk on land. These grebes may be confused with the
least grebe The least grebe (''Tachybaptus dominicus''), an aquatic bird, is the smallest member of the grebe family. It occurs in the New World from the southwestern United States and Mexico to Argentina, and also on Trinidad and Tobago, the Bahamas and the ...
, although that species is much smaller and has a thinner bill. Other similarly sized grebes are very distinct in plumage, i.e. the
eared grebe The black-necked grebe or eared grebe (''Podiceps nigricollis'') is a member of the grebe family of water birds. It was described in 1831 by Christian Ludwig Brehm. There are currently three accepted subspecies, including the nominate subspeci ...
and
horned grebe The horned grebe or Slavonian grebe (''Podiceps auritus'') is a relatively small waterbird in the family Podicipedidae. There are two known subspecies: ''P. a. auritus'', which breeds in the Palearctic, and ''P. a. cornutus'', which breeds in ...
. Both species bear much more colorful breeding plumage, with rufous sides, golden crests along the side of the head against contrasting slaty color (also a rufous neck in the horned); while in winter, both the eared and horned grebes are pied with slaty and cream color and have red eyes. Because of the pied-billed grebe's duck-like habits, some inexperienced observers may confuse it with a duck. However, pied-billed grebes have a very different bill shape (shorter, pointed at the tip, and flattened along the sides), as well as being shorter-necked and shorter-bodied than a duck. Also, unlike ducks, the closest living relatives of the grebe family are
flamingos Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...
. Pied-billed grebes are part of the order
Saurischia Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithis ...
.


Vocalization

Its call is unique, loud and sounds like a "whooping ''kuk-kuk-cow-cow-cow-cowp-cowp''." Its call is similar to the
yellow-billed cuckoo The yellow-billed cuckoo (''Coccyzus americanus'') is a cuckoo. Common folk-names for this bird in the southern United States are rain crow and storm crow. These likely refer to the bird's habit of calling on hot days, often presaging rain or th ...
.


Distribution and habitat

They are most commonly found throughout
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
year round. During the summer breeding season, they are most prevalent in central, northern and northeastern
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
. If they live in an area where the water freezes in the winter they will migrate. Migrating birds generally meet with year-round birds in September and October. They migrate at night. Most migratory birds leave in March or April. They make occasional appearances in Europe and
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
. In the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
, pied-billed grebe visits have numbered 45 sightings as of 2019, appearing generally in October to January. One bird in England bred with a
little grebe The little grebe (''Tachybaptus ruficollis''), also known as dabchick, is a member of the grebe family of water birds. The genus name is from Ancient Greek ''takhus'' "fast" and ''bapto'' "to sink under". The specific ''ruficollis'' is from Latin ...
, producing
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
young. It is the only grebe on record to have visited the Galapagos Islands. Pied-billed grebes are found in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
with emergent vegetation, such as
cattails ''Typha'' is a genus of about 30 species of monocotyledonous flowering plants in the family Typhaceae. These plants have a variety of common names, in British English as bulrush or reedmace, in American English as reed, cattail, or punks, in ...
. They are occasionally found in
salt water Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
. When breeding they are found in emergent vegetation near open water, and in the winter they are primarily found in open water due to the lack of nests to maintain. They may live near rivers, but prefer still water. They may be found in higher elevations when migrating. They will breed in restored and man-made wetlands. Pied-billed grebes live approximately 10–12 years.


Behaviour

Pied-billed grebes rarely fly. They make a slow dive frequently, especially when in danger, diving to about or less. They dive for about 30 seconds and may move to a more secluded area of the water, allowing only the head to be visible to watch the danger dissipate. This frequency in diving has earned them the description of being reclusive or shy in nature. It has also earned them nicknames like "hell-diver." Pied-billed grebes can trap air in their feathers, which controls their buoyancy. They rarely spend time in flocks. Their
courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
include calling and sometimes duets. Males will show territorial behaviour if another male is at the edge of his territory. They face each other and then turn their heads and bills up. Then they turn away and start calling. Then they turn back around to look at one another.


Breeding

The pied-billed grebe breeds in south-central
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, throughout the United States,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, and temperate
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. These grebes may lay up to two sets of eggs a year. Their nests sit on top of the water, their eggs sitting in vegetation that resides in the water. Grebes lay between three and ten bluish white smooth elliptical eggs with the female starting the incubation process. They are incubated for around 23 days by both parents, with the female taking over incubation duties towards the end of that time period. They will cover the nest with nesting material if they have to leave it for an extended period of time. Young grebes may leave the nest within one day of hatching. They are
downy Downy, also known as Lenor in Europe, Russia and Japan, is a brand of fabric softener produced by Procter & Gamble that was introduced in 1960. Lenor is a brand name of fabric softener and dryer sheets, also produced by P&G, sold in Europe, Rus ...
at birth. Yellow skin is seen between the
lore Lore may refer to: * Folklore, acquired knowledge or traditional beliefs * Oral lore or oral tradition, orally conveyed cultural knowledge and traditions Places * Loré, former French commune * Loré (East Timor), a city and subdistrict in Lau ...
and top of the head. They do not swim well and stay out of the water. They sleep on their parents' backs. Within four weeks they start swimming. When alerted they will climb on the back of a parent grebe and eventually mature to dive under the water like their parents. Both parents share the role of raising the young – both feeding and carrying them on their backs. Sometimes the parents will dive underwater to get food with the chicks on their backs.


Diet

Pied-billed grebes feed mainly on aquatic
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s, and also on small
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
and
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s (
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s,
tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found i ...
s). They dive to obtain food. Their bills allow them to crush
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s, like crawfish. They may also eat plants. They have been shown to eat their own feathers, like other grebes, to aid in digestion (prevent injury from small bones).Ehrlich, Paul; David Dobkin, and Darryl Wheye (1988). ''The Birder's Handbook''. New York: Simon and Schuster. They will also feed their feathers to their young.


Threats

They are extremely sensitive to disturbances, especially by humans. While breeding, if scared, adults may abandon their nests without protecting the eggs. The waves from
boats A boat is a watercraft of a large range of types and sizes, but generally smaller than a ship, which is distinguished by its larger size, shape, cargo or passenger capacity, or its ability to carry boats. Small boats are typically found on inl ...
can destroy the nests and their sounds easily frighten the birds.


In culture

Pied-billed grebe feathers are thick and soft. Their feathers were formerly used as decorations on hats and
earmuffs Earmuffs are clothing accessories or personal protective equipment designed to cover a person's ears for hearing protection or warmth. They consist of a thermoplastic or metal head-band that fits over the top or back of the head, and a cushion ...
and they were hunted in the eastern United States, in the 19th century.


Status

The grebes are declining in
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
. The reasons are unknown. The states of
Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York to the west, and Long Island Sound to the south. Its cap ...
and
New Hampshire New Hampshire is a U.S. state, state in the New England region of the northeastern United States. It is bordered by Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...
have declared the pied-billed grebe as
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
. In
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...
and
Massachusetts Massachusetts (Massachusett language, Massachusett: ''Muhsachuweesut assachusett writing systems, məhswatʃəwiːsət'' English: , ), officially the Commonwealth of Massachusetts, is the most populous U.S. state, state in the New England ...
, they have been declared
threatened Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensat ...
. In
Vermont Vermont () is a state in the northeast New England region of the United States. Vermont is bordered by the states of Massachusetts to the south, New Hampshire to the east, and New York to the west, and the Canadian province of Quebec to ...
they are of "special concern." In
Rhode Island Rhode Island (, like ''road'') is a U.S. state, state in the New England region of the Northeastern United States. It is the List of U.S. states by area, smallest U.S. state by area and the List of states and territories of the United States ...
they are
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
. Pied-billed grebes are protected under the
Migratory Bird Treaty Act of 1918 The Migratory Bird Treaty Act of 1918 (MBTA), codified at (although §709 is omitted), is a United States federal law, first enacted in 1918 to implement the convention for the protection of migratory birds between the United States and Canada . ...
.
Habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
is the grebe's biggest threat. The draining, filling, and general destruction of wetlands causes a loss in their breeding habitats. However, they are still common in the majority of their distribution areas.


References


External links

* *
Pied-billed Grebe Species Account
– Cornell Lab of Ornithology {{Authority control
pied-billed grebe The pied-billed grebe (''Podilymbus podiceps'') is a species of the grebe family of water birds. Because the Atitlán grebe (''Podilymbus gigas'') has become extinct, the Pied-Billed Grebe is now the sole extant member of the genus ''Podilymbus'' ...
Birds of the Americas
pied-billed grebe The pied-billed grebe (''Podilymbus podiceps'') is a species of the grebe family of water birds. Because the Atitlán grebe (''Podilymbus gigas'') has become extinct, the Pied-Billed Grebe is now the sole extant member of the genus ''Podilymbus'' ...
Taxa named by Carl Linnaeus