Picea jezoensis
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''Picea jezoensis'' (sometimes misspelled ''Picea yezoensis''), the dark-bark spruce, Ezo spruce, Yezo spruce, or Jezo spruce, is a large
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to 30–50 m tall and with a trunk diameter of up to 2 m. It is native to northeast
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, from the mountains of central
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
and the
Changbai Mountains The Changbai Mountains ( simplified Chinese:长白山; traditional Chinese:長白山) are a major mountain range in Northeast Asia that extends from the Northeast Chinese provinces of Heilongjiang, Jilin and Liaoning, across the border bet ...
on the
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
-
North Korea North Korea, officially the Democratic People's Republic of Korea (DPRK), is a country in East Asia. It constitutes the northern half of the Korean Peninsula and shares borders with China and Russia to the north, at the Yalu (Amnok) and T ...
border, north to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
, including the
Sikhote-Alin The Sikhote-Alin (russian: Сихотэ́-Али́нь, , , ) is a mountain range in Primorsky and Khabarovsk Krais, Russia, extending about to the northeast of the Russian Pacific seaport of Vladivostok. The highest summits are Tordoki Yani a ...
,
Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese language, Japanese: or ) are a volcanic archipelago currently administered as part of Sakh ...
,
Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh ...
and
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
. It is found in cold but humid
temperate rain forests Temperate rainforests are coniferous or broadleaf forests that occur in the temperate zone and receive heavy rain. Temperate rain forests occur in oceanic moist regions around the world: the Pacific temperate rain forests of North American Pac ...
, and nowhere does its range extend more than 400 km from the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the conti ...
. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''jezoensis'' derives from
Ezo (also spelled Yezo or Yeso) is the Japanese term historically used to refer to the lands to the north of the Japanese island of Honshu. It included the northern Japanese island of Hokkaido, which changed its name from "Ezo" to "Hokkaidō" in 18 ...
, an old name for
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
and other islands north of the Japanese island of
Honshu , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island s ...
, where the species is found. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
is thin and scaly, becoming fissured in old trees. The crown is broad conic. The shoots are pale buff-brown, glabrous (hairless) but with prominent pulvini. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are needle-like, 15–20 mm long, 2 mm broad, flattened in cross-section, dark green above with no
stomata In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is b ...
, and blue-white to white below with two dense bands of stomata. The cones are pendulous, slender cylindrical, 4–7 cm long and 2 cm broad when closed, opening to 3 cm broad. They have thin, flexible scales 12–18 mm long. They are green or reddish, maturing pale brown 5–6 months after pollination. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s are black, 3 mm long, with a slender, 6–8 mm long pale brown wing. There are two geographical
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
, treated as varieties by some authors, and as distinct species by others: *''Picea jezoensis'' subsp. ''jezoensis'' (Ezo spruce). All of the range except as below, south to
Hokkaidō is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
, Japan. Shoots very pale buff-brown, almost white; stomatal bands blue-white; cones pale brown with flexible scales. *''Picea jezoensis'' subsp. ''hondoensis'' (Mayr) P. A. Schmidt (Hondo spruce). An isolated southern population on high mountains in central
Honshū , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island sepa ...
, Japan. Shoots buff-brown to orange-brown, less often very pale; stomatal bands bright white; cones orange-brown with stiffer scales. Ezo spruce is very closely related to Sitka spruce (''
Picea sitchensis ''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to almost tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-larg ...
''), which replaces it on the opposite side of the north Pacific. They, particularly subsp. ''jezoensis'', can be difficult to distinguish, with the absence of stomata on the upper surface of the leaves of ''P. jezoensis'' being the best feature. Its leaves are also somewhat blunter, less sharply spine-tipped, than Sitka Spruce.


Usage

Jezo spruce is important in the Russian Far East and northern Japan, for timber and paper production. Much of what is cut is harvested unsustainably (and often illegally) from pristine natural
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s. It is also occasionally planted as an ornamental tree in large gardens. The
Ainu Ainu or Aynu may refer to: *Ainu people, an East Asian ethnic group of Japan and the Russian Far East *Ainu languages, a family of languages **Ainu language of Hokkaido **Kuril Ainu language, extinct language of the Kuril Islands **Sakhalin Ainu la ...
string instrument called '' tonkori'' has a body made from Jezo Spruce.


References


External links


Arboretum de Villardebelle - photos of cones of ''Picea jezoensis'' and related spruces
{{Authority control jezoensis Trees of China Trees of Japan Trees of Korea Trees of Siberia Trees of continental subarctic climate Ornamental trees Least concern plants