Petrels
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Petrels are tube-nosed
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s in the
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
order
Procellariiformes Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are oft ...
.


Description

The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pac ...
family, Diomedeidae). Having a
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
record that was assumed to extend back at least 60 million years, the Procellariiformes was long considered to be among the older bird groupings, other than the
ratite A ratite () is any of a diverse group of flightless, large, long-necked, and long-legged birds of the infraclass Palaeognathae. Kiwi, the exception, are much smaller and shorter-legged and are the only nocturnal extant ratites. The systematics ...
s, with presumably distant ties to
penguin Penguins (order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adap ...
s and
loon Loons ( North American English) or divers ( British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order ...
s. However, recent research and fossil finds such as ''
Vegavis ''Vegavis'' is a genus of extinct bird that lived during the Late Cretaceous (Maastrichtian stage) of Antarctica, some 68 to 66 mya. Among modern birds, most studies show that ''Vegavis'' is most closely related to ducks and geese (Anatidae), b ...
'' show that the
Galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
(
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera native range is restricted to Eurasia ...
s,
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
and relatives), and
Anseriformes Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which in ...
(
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s,
geese A goose ( : geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera '' Anser'' (the grey geese and white geese) and ''Branta'' (the black geese). Some other birds, mostly related to the she ...
) are still not fully resolved.


Known species

All the members of the order are exclusively
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
in distribution—returning to land only to breed. The family
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which als ...
is the main radiation of medium-sized true petrels, characterised by united nostrils with medium septum, and a long outer functional primary feather. It is dominant in the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smal ...
s, but not so in the Northern Hemisphere. It includes a number of petrel groups, the relationships between which have finally been resolved to satisfaction. * The fulmarine petrels: seven species of surface predators and
filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
s, breed in high latitudes but migrate along cool currents to the north. All but ''Fulmarus'' are essentially confined to the south, ''Fulmarus'' apparently colonised the Northern Hemisphere during the Early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
. ** The huge
giant petrel Giant petrels form a genus, ''Macronectes'', from the family Procellariidae, which consists of two species. They are the largest birds of this family. Both species are restricted to the Southern Hemisphere, and though their distributions overla ...
s, genus ''Macronectes'', which are convergent with the albatrosses ** The true
fulmar The fulmars are tubenosed seabirds of the family Procellariidae. The family consists of two extant species and two extinct fossil species from the Miocene. Fulmars superficially resemble gulls, but are readily distinguished by their flight on ...
s, genus ''Fulmarus'' **
Antarctic petrel The Antarctic petrel (''Thalassoica antarctica'') is a boldly marked dark brown and white petrel, found in Antarctica, most commonly in the Ross and Weddell Seas. They eat Antarctic krill, fish, and small squid. They feed while swimming but c ...
''Thalassoica antarctica'' ** Cape petrel ''Daption capense'' **
Snow petrel The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that has been seen at the Geographic South Pole, along with the Antarctic petrel and the south polar skua, which have the most s ...
''Pagodroma nivea'' * The
prions Prions are misfolded proteins that have the ability to transmit their misfolded shape onto normal variants of the same protein. They characterize several fatal and transmissible neurodegenerative diseases in humans and many other animals. It i ...
: A specialised group of a few very numerous species, all southern. They have a small, fulmar-like form and mostly filter-feed on zooplankton. ** '' Pachyptila'', the prions proper * The procellariine petrels, larger or mid-sized species feeding on fish and molluscs which are fairly close to the prions: ** ''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The specta ...
'' ** '' Bulweria'' *
Shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae. They have a global marine distribution, but are most common in temperate and cold waters, and are pelagic outside the breeding season. Description These tub ...
s: There are numerous species in several genera with a medium number of species. ** ''
Calonectris ''Calonectris'' is a genus of seabirds. The genus name comes from Ancient Greek ''kalos'', "good" and ''nectris'', "swimmer". The genus comprises four large shearwaters. There are two other shearwater genera. ''Puffinus'', which comprises abou ...
'' ** ''
Puffinus ''Puffinus'' is a genus of seabirds in the order Procellariiformes that contains about 20 small to medium-sized shearwaters. Two other shearwater genera are named: '' Calonectris'', which comprises three or four large shearwaters, and '' Ardenna' ...
'', which is two rather distinct groups of larger and smaller species ** ''
Pseudobulweria __NOTOC__ ''Pseudobulweria'' is a genus of seabirds in the family Procellariidae. They have long been retained with the gadfly petrel genus ''Pterodroma'' despite morphological differences. Mitochondrial DNA cytochrome ''b'' sequence analysis ...
'' **
Kerguelen petrel The Kerguelen petrel (''Aphrodroma brevirostris'') is a small (36 cm long) slate-grey seabird in the family Procellariidae. It is the only species placed in the genus ''Aphrodroma''. It is a pelagic, circumpolar seabird of the Southern Oc ...
''Lugensa brevirostris'' * The
gadfly petrel The gadfly petrels or ''Pterodroma'' are a genus of about 35 species of petrels, part of the seabird order Procellariiformes. The gadfly petrels are named for their speedy weaving flight, as if evading gadflies ( horseflies). The flight action is ...
s: These are a considerable number of agile short-billed petrels in the genus ''Pterodroma'' which include the endangered
Bermuda petrel The Bermuda petrel (''Pterodroma cahow'') is a gadfly petrel. Commonly known in Bermuda as the cahow, a name derived from its eerie cries, this nocturnal ground-nesting seabird is the national bird of Bermuda and can be found pictured on Bermudia ...
or cahow and a considerable number of forms rendered
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
by human activity. The families
Oceanitidae Austral storm petrels, or southern storm petrels, are seabirds in the family Oceanitidae, part of the order Procellariiformes. These smallest of seabirds feed on planktonic crustaceans and small fish picked from the surface, typically while hove ...
and Hydrobatidae are the storm petrels, small pelagic petrels with a fluttering flight which often follow ships. The family Pelecanoididae is the four species of diving petrels, genus ''Pelacanoides''. These are
auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...
-like small petrels of the southern oceans. The word ''petrel'' (first recorded in that spelling 1703) comes from earlier (''ca.'' 1670) ''pitteral''; the English explorer
William Dampier William Dampier (baptised 5 September 1651; died March 1715) was an English explorer, pirate, privateer, navigator, and naturalist who became the first Englishman to explore parts of what is today Australia, and the first person to circumnav ...
wrote the bird was so called from its way of flying with its feet just skimming the surface of the water, recalling
Saint Peter ) (Simeon, Simon) , birth_date = , birth_place = Bethsaida, Gaulanitis, Syria, Roman Empire , death_date = Between AD 64–68 , death_place = probably Vatican Hill, Rome, Italia, Roman Empire , parents = John (or Jonah; Jona) , occupat ...
's walk on the sea of Galilee (Matthew xiv.28); if so, it likely was formed in English as a diminutive of Peter (< Old french: Peterelle (?) > Late lat, Peterellus < Late lat, Petrus < grc-gre, Πέτρος, Petros < grc-gre, πέτρα, petra = "stone").


See also

*
Skua The skuas are a group of predatory seabirds with seven species forming the genus ''Stercorarius'', the only genus in the family Stercorariidae. The three smaller skuas, the long-tailed skua, the Arctic skua, and the pomarine skua are calle ...
*
Agreement on the Conservation of Albatrosses and Petrels The Agreement on the Conservation of Albatrosses and Petrels (ACAP) is a legally binding international agreement signed in 2001 and entered into force on 1 February 2004 when South Africa ratified as the fifth Party to the Agreement. It was crea ...


References


External links

*
Petrel and shearwater videos
on the Internet Bird Collection
Agreement on the Conservation of Albatrosses and Petrels (ACAP)
__NOTOC__ {{Authority control Procellariiformes Seabirds Bird common names