Pellet (ornithology)
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A pellet, in
ornithology Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
, is the mass of undigested parts of a bird's food that some bird species occasionally regurgitate. The contents of a bird's pellet depend on its diet, but can include the
exoskeleton An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the ...
s of insects, indigestible plant matter, bones, fur, feathers, bills, claws, and teeth. In falconry, the pellet is called a ''casting''. The passing of pellets allows a bird to remove indigestible material from its proventriculus, or glandular
stomach The stomach is a muscular, hollow organ in the gastrointestinal tract of humans and many other animals, including several invertebrates. The stomach has a dilated structure and functions as a vital organ in the digestive system. The stomach i ...
. In birds of
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
, the regurgitation of pellets serves the bird's health in another way, by "scouring" parts of the digestive tract, including the
gullet The esophagus (American English) or oesophagus (British English; both ), non-technically known also as the food pipe or gullet, is an organ in vertebrates through which food passes, aided by peristaltic contractions, from the pharynx to the ...
. Pellets are formed within six to ten hours of a meal in the bird's
gizzard The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including archosaurs (pterosaurs, crocodiles, alligators, dinosaurs, birds), earthworms, some gastropods, so ...
(muscular stomach). Ornithologists may collect one species' pellets over time to analyze the seasonal variation in its eating habits. One advantage of collecting pellets is that it allows for the determination of diet without the killing and dissection of the bird. Pellets are found in different locations, depending on the species. In general, these are roosting and nesting sites: for most hawks and owls, under coniferous trees; for barn owls, at the bases of cliffs or in barns and silos; for yet other species of owls, at their burrows or in marsh and field grasses.Terres, John. K. ''The Audubon Society Encyclopedia of North American Birds'', New York: Alfred A. Knopf, 1980. Hawk and
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
pellets are grey or brown, and range in shape from spherical to oblong or plug-shaped. In large birds, they are one to two inches long, and in
songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 5000 ...
s, about half an inch. Many other species produce pellets, including grebes, herons, cormorants,
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s,
tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists of e ...
s,
kingfisher Kingfishers are a family, the Alcedinidae, of small to medium-sized, brightly colored birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species found in the tropical regions of Africa, Asia, and Oceania, ...
s,
crow A crow is a bird of the genus '' Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifica ...
s, jays,
dipper Dippers are members of the genus ''Cinclus'' in the bird family Cinclidae, so-called because of their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater. Taxonomy The genus ''Cinclus'' ...
s, shrikes,
swallow The swallows, martins, and saw-wings, or Hirundinidae, are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...
s, and most
shorebird 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s. Ornithologists examining pellets have discovered unusual items in them—even bird bands that were once attached to a smaller species that was consumed by the predator bird. In the United States,
screech owl Screech owls are typical owls (Strigidae) belonging to the genus ''Megascops'' with 23 living species. For most of the 20th century, this genus was merged with the Old World scops owls in ''Otus'', but nowadays it is again considered separate ...
pellets have contained bands from a tufted titmouse,
black-capped chickadee The black-capped chickadee (''Poecile atricapillus'') is a small, non-migratory, North American songbird that lives in deciduous and mixed forests. It is a passerine bird in the tit family, the Paridae. It is the state bird of Massachusetts and ...
, and
American goldfinch The American goldfinch (''Spinus tristis'') is a small North American bird in the finch family. It is migratory, ranging from mid-Alberta to North Carolina during the breeding season, and from just south of the Canada–United States border t ...
. In 1966, a golden eagle pellet in
Oregon Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idaho. T ...
was found to contain a band placed on an
American wigeon The American wigeon (''Mareca americana''), also known as the baldpate, is a species of dabbling duck found in North America. Formerly assigned to ''Anas'', this species is classified with the other wigeons in the dabbling duck genus '' Mareca'' ...
four months earlier, and away in southern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. The hair, bones and other body parts (such as limbs, skin fragments, and even faeces) of rodents found in owl pellets may carry viable rodent
viruses A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsky's ...
and
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of prokaryotic microorganisms. Typically a few micrometr ...
. for this reason, they are sometimes sterilized before study. Smith ''et al.'' described two pellet-borne outbreaks of ''
Salmonella typhimurium ''Salmonella enterica'' subsp. ''enterica'' is a subspecies of ''Salmonella enterica'', the rod-shaped, flagellated, aerobic, Gram-negative bacterium. Many of the pathogenic serovars of the ''S. enterica'' species are in this subspecies, includin ...
'' in schools where unsterile pellets were dissected. Rodents tend to avoid owl pellets, apparently due to their infective potential.


In falconry

In falconry, casting involves giving a hawk roughage, which is regurgitated later to purge and cleanse its
crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydropon ...
. Hawks in the wild will produce castings of their own accord, in the form of small pellets of fur, feathers, and other indigestible material, regurgitated hours after it has eaten. Of the roughage used for falcons in captivity, there are two kinds: plumage, and cotton; the latter of which is generally in pellets about the size of hazelnuts, made of soft fine cotton, and conveyed into the hawk's gorge after supper. In the morning, the hawk will have ''cast'' them out, at which time they should be examined for color and condition to determine the state of the hawk's body. If they are cast out round, white, without stench, nor very moist, then the hawk is healthy; if otherwise, particularly black, green, slimy, or the like, the hawk is ill. The casting of plumage is examined in the same manner. The yellow
down feathers The down of birds is a layer of fine feathers found under the tougher exterior feathers. Very young birds are clad only in down. Powder down is a specialized type of down found only in a few groups of birds. Down is a fine thermal insulator an ...
of day-old
rooster The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
chicks, which are a common diet among hawks in captivity, can be enough to induce casting. However, rabbit fur and feathers from game birds are better suited for this purpose.Ford, Emma. ''Falconry: Art and Practice, Revised Edition''. Octopus Publishing Group - Cassell Illustrated. 1998. . p 23. Casting will occur at the same time daily, provided that hawks are flown at the same time daily. Hawks in captivity are usually weighed immediately before being flown. If they were fed roughage the previous day, they should cast it out before being weighed, to avoid an inaccurate measurement. Casting is a necessary part of a hawk's survival. Not only does it clear out the crop, but the roughage that is regurgitated cleans bacteria from the walls of the crop. If hawks in captivity are not provided with this necessary roughage, and instead are simply fed meat, they will become seriously ill and unreleasable into the wild.Elsberry, Wesley R.
Falconry Jargon
. ''Austringer''. URL accessed 2006-05-30.


References


External links

{{commons category, Bird pellets

Bird feeding