Paraptenodytes
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''Paraptenodytes'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
penguin Penguins (order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adap ...
s which contains two or three
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
sized between a Magellanic penguin and an
emperor penguin The emperor penguin (''Aptenodytes forsteri'') is the tallest and heaviest of all living penguin species and is endemic to Antarctica. The male and female are similar in plumage and size, reaching in length and weighing from . Feathers of t ...
(''Aptenodytes forsteri''). They are known from
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
bones ranging from a partial skeleton and some additional material in the case of ''P. antarcticus'', the type specimen for the genus, and a single
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a r ...
in the case of ''P. brodkorbi''. The latter species is therefore often considered invalid; a recent studyBertelli ''et al.'' (2006) considers it indeed valid, but distinct enough not to belong into ''Paraptenodytes''. The fossils were found in the Santa Cruz and
Chubut Province Chubut ( es, Provincia del Chubut, ; cy, Talaith Chubut) is a province in southern Argentina, situated between the 42nd parallel south (the border with Río Negro Province), the 46th parallel south (bordering Santa Cruz Province), the Ande ...
s of
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...
, in the Gaiman, Monte León and Santa Cruz Formations of Early to Middle Miocene age.''Paraptenodytes''
at
Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was crea ...
.org
Later occurrences are apparently from Late Miocene or possibly even Early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Arthrodytes'', they form the subfamily Paraptenodytinae, which is not an ancestor of modern penguins.


References


Bibliography

* Bertelli, Sara; Giannini, Norberto P. & Ksepka, Daniel T. (2006): Redescription and Phylogenetic Position of the Early Miocene Penguin ''Paraptenodytes antarcticus'' from Patagonia. ''American Museum Novitates'' 3525: 1-36. DOI: 10.1206/0003-0082(2006)3525 :RAPPOT.0.CO;2PDF fulltext
* Stucchi, Marcelo; Urbina, Mario & Giraldo, Alfredo (2003): Una nueva especie de Spheniscidae del Mioceno Tardío de la Formación Pisco, Perú. ''Bulletin Institut Français d'Études Andines'' 32(2): 361–375
PDF fulltext


Further reading

* Ameghino, Florentino (1891): Enumeración de las aves fósiles de la Repúiblica Argentina. ''Revista Argentina de Historia Natural'' 1: 441–445. * Simpson, George Gaylord (1946): Fossil penguins. ''Bulletin of the American Museum of Natural History'' 87: 7-99
PDF fulltext
* Simpson, George Gaylord (1971): Conspectus of Patagonian fossil penguins. ''American Museum Novitates'' 2488: 1-37
PDF fulltext
{{Taxonbar, from=Q3895277 Bird genera Extinct penguins Miocene birds of South America Pliocene birds of South America Laventan Colloncuran Friasian Santacrucian Colhuehuapian Neogene Argentina Fossils of Argentina Gaiman Formation Fossil taxa described in 1891 Taxa named by Florentino Ameghino