''Papilio memnon'', the great Mormon, is a large

butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group compris ...

native to southern Asia that belongs to the swallowtail

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

. It is widely distributed and has thirteen

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...

. The female is polymorphic and with mimetic forms.

Range

Its range includes north-eastern India (including

Sikkim Sikkim (; ) is a state in Northeastern India. It borders the Tibet Autonomous Region of China in the north and northeast, Bhutan in the east, Province No. 1 of Nepal in the west and West Bengal in the south. Sikkim is also close to the Sil ...

Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...

and

Nagaland Nagaland () is a landlocked state in the northeastern region of India. It is bordered by the Indian states of Arunachal Pradesh to the north, Assam to the west, Manipur to the south and the Sagaing Region of Myanmar to the east. Its capital cit ...

Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...

Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...

Nicobar Islands The Nicobar Islands are an archipelagic island chain in the eastern Indian Ocean. They are located in Southeast Asia, northwest of Aceh on Sumatra, and separated from Thailand to the east by the Andaman Sea. Located southeast of the Indian ...

Andaman Islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between t ...

(stragglers only), western, southern and eastern China (including

Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...

, southern

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...

including

Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yon ...

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...

Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist s ...

Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...

, Kampuchea,

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federal constitutional monarchy consists of thirteen states and three federal territories, separated by the South China Sea into two regions: Peninsular Malaysia and Borneo's East Mal ...

and

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Gui ...

(

Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...

Mentawai Islands Mentawai may refer to: * Mentawai Islands * Mentawai Strait * Mentawai people * Mentawai language The Mentawai language is an Austronesian language, spoken by the Mentawai people of the Mentawai Islands, West Sumatra, Indonesia. Dialects Ac ...

Nias Nias ( id, Pulau Nias, Nias language: ''Tanö Niha'') (sometimes called Little Sumatra in English) is an island located off the western coast of Sumatra, Indonesia. Nias is also the name of the archipelago () of which the island is the centre ...

, Batu,

Simeulue Simeulue is an island of Indonesia, off the west coast of Sumatra. It covers an area of 1754 square kilometres (677 square miles), including minor offshore islands. It had a population of 80,674 at the 2010 census and 92,865 at the 2020 census ...

, Bangka,

Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...

Kalimantan Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo. In 2019, ...

and the

Lesser Sunda Islands The Lesser Sunda Islands or nowadays known as Nusa Tenggara Islands ( id, Kepulauan Nusa Tenggara, formerly ) are an archipelago in Maritime Southeast Asia, north of Australia. Together with the Greater Sunda Islands to the west they make up ...

Status

This species is common and not threatened. The cultivation of

citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...

all over Southern Asia provides an abundance of food plants.

Description and polymorphy

The butterfly is large with a span. It has four male and many female forms, the females being highly polymorphic and many of them being mimics of unpalatable butterflies. This species has been studied extensively for understanding the genetic basis for polymorphy and

Batesian mimicry Batesian mimicry is a form of mimicry where a harmless species has evolved to imitate the warning signals of a harmful species directed at a predator of them both. It is named after the English naturalist Henry Walter Bates, after his work on ...

. As many as twenty-six female forms are reported.

Typical form ''agenor''

*Male. Tailless, above deep blue to black. It may or may not have red streak on the forewing at the base of the cell. *Female. Tailless. Upperside forewing ground colour

sepia Sepia may refer to: Biology * ''Sepia'' (genus), a genus of cuttlefish Color * Sepia (color), a reddish-brown color * Sepia tone, a photography technique Music * ''Sepia'', a 2001 album by Coco Mbassi * ''Sepia'' (album) by Yu Takahashi * " ...

, streaked with greyish white. The basal third part of the cell is red and is touched outwardly with white. Upper hindwing is blue black. It has five to seven yellow or white discal patches.

Female form ''butlerianus''

*Tailless. Resembles the typical male. Both wings are dark sepia. The forewing has a white area on the inner margin. The hindwing is scaled with blue.

Female form ''alcanor''

*Tailed. The sides of the abdomen are yellow. *Upperside forewing greyish brown with veins and streaks between them black. The cell is red at the base. There is a velvety black patch at the bases of veins 1 and 2 of the upperside forewing. *Upperside hindwing is black with part of the cell white. There are white streaks around it. The tornus is red with a large black spot. There is a row of red terminal spots between the vein.

Male and female form ''polymnestoroides''

*Tailless. *Male. Upperside hindwing and forewing have short blue discal stripes. *Female. The upperside forewing is sepia with pale grey streaks amongst the veins. The base of the cell is red. The upperside hindwing is velvety brown with a blue discal area and has black spots, as in the case of the blue Mormon (''P. polymnestor''), which it mimics.

Habitat

This species flies up to in the Himalayas, but is most common at low elevations.

Behaviour

This butterfly is found in forest clearings. It is very common and is also seen amongst human habitation. It visits flowers of ''

Poinsettia The poinsettia ( or ) (''Euphorbia pulcherrima'') is a commercially important flowering plant species of the diverse spurge family Euphorbiaceae. Indigenous to Mexico and Central America, the poinsettia was first described by Europeans in 183 ...

'', ''

Jasminum Jasmine ( taxonomic name: ''Jasminum''; , ) is a genus of shrubs and vines in the olive family (Oleaceae). It contains around 200 species native to tropical and warm temperate regions of Eurasia, Africa, and Oceania. Jasmines are widely cultiva ...

'', ''

Lantana ''Lantana'' () is a genus of about 150 species of perennial flowering plants in the verbena family, Verbenaceae. They are native to tropical regions of the Americas and Africa but exist as an introduced species in numerous areas, especially in ...

'', '' Canna'' and ''

Salvia ''Salvia'' () is the largest genus of plants in the sage family Lamiaceae, with nearly 1000 species of shrubs, herbaceous perennials, and annuals. Within the Lamiaceae, ''Salvia'' is part of the tribe Mentheae within the subfamily Nepetoide ...

''. It usually flies above the ground. The butterfly is known to

mud-puddle Mud-puddling, or simply puddling, is a behaviour most conspicuous in butterflies, but occurs in other animals as well, mainly insects; they seek out nutrients in certain moist substances such as rotting plant matter, mud and carrion and they suc ...

. The males are much more common than females. The female forms ''butlerianus'' and ''alcanor'' are especially uncommon.

Life cycle

The larva resembles that of the

common Mormon ''Papilio polytes'', the common Mormon, is a common species of swallowtail butterfly widely distributed across Asia. This butterfly is known for the mimicry displayed by the numerous forms of its females which mimic inedible red-bodied swallo ...

(''P. polytes''), being green with whitish markings. It is heavily parasitised.

Gallery

Great Mormon, dorsal view.jpg, Male ''agenor'' Papilio memnon-agenor female.jpg, Female ''agenor'' Papilio memnon-alcanor female dorsal.jpg, Female ''alcanor'' upperside Papilio memnon-alcanor female ventral.jpg, Female ''alcanor'' underside Close wing Nectaring of Papilio memnon Linnaeus, 1758 – Great Mormon (Female) Form butlerianus WLB DSC 9451.jpg, Female ''butlerianus'' underside Open wing nectaring position of Papilio memnon Linnaeus, 1758 – Great Mormon (Female) Form butlerianusDSC 2479.jpg, Female ''butlerianus'' upperside Papilio memnon Linne.jpg, Male Underside Great mormons mating.JPG, Mating pair Papilio memnon heronus larva 20140614.jpg, Larva

References

External links

* {{Taxonbar, from=Q1303868

memnon In Greek mythology, Memnon (; Ancient Greek: Μέμνων means 'resolute') was a king of Aethiopia and son of Tithonus and Eos. As a warrior he was considered to be almost Achilles' equal in skill. During the Trojan War, he brought an army ...

Butterflies of Oceania Butterflies of Indochina Butterflies of Asia Butterflies of Indonesia Butterflies of Malaysia Butterflies of Singapore Lepidoptera of the Philippines Lepidoptera of New Guinea Butterflies described in 1758 Taxa named by Carl Linnaeus