Pandaka pygmaea
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The dwarf pygmy goby or Philippine goby (''Pandaka pygmaea'') is a
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
species of fish in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Gobionellinae The Gobionellinae are a subfamily of fish which was formerly classified in the family Gobiidae, the gobies, but the 5th Edition of '' Fishes of the World'' classifies the subfamily as part of the family Oxudercidae. Members of Gobionellinae most ...
from
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estua ...
and
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
areas in Southeast Asia. It is one of the smallest fish species in the world. Males reach maturity at a
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of and can reach up to in standard length, while the females can grow up to in
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
. Adults weigh around . It is known as ''bia'' and ''tabios'' in the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
., ''Guide to the Philippine Flora and Fauna, Band IX'', (undated).


Distribution and habitat

''P. pygmaea'' was initially reported as being
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Malabon Malabon, officially the City of Malabon ( fil, Lungsod ng Malabon), is a 1st class highly urbanized city in the National Capital Region of the Philippines. According to the 2020 census, it has a population of 380,522 people. Located just north ...
,
Metro Manila Metropolitan Manila (often shortened as Metro Manila; fil, Kalakhang Maynila), officially the National Capital Region (NCR; fil, link=no, Pambansang Punong Rehiyon), is the capital region, seat of government and one of three List of metrop ...
in the Philippines, where found at shady river banks. The species has now been
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
from this area as it was
reclaimed Land reclamation, usually known as reclamation, and also known as land fill (not to be confused with a waste landfill), is the process of creating new Terrestrial ecoregion, land from oceans, list of seas, seas, Stream bed, riverbeds or lak ...
, leading to its status as critically endangered by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. More recently, it has been discovered at a range of other locations in Southeast Asia, including
Culion Island Culion, officially the Municipality of Culion ( tgl, Bayan ng Culion), is a 3rd class municipality in the province of Palawan, Philippines. According to the 2020 census, it has a population of 23,213 people. Culion consists primarily of Culion ...
,
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
,
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
and
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
. As part of the aquarium trade, it was imported into Germany in 1958. ''P. pygmaea'' lives in shallow, tropical
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estua ...
and
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
areas where mainly found on muddy bottoms or among plants. Also known before as National Fish in the Philippines on 1994 until recently they changed it to milk fish.


Appearance and anatomy

A colorless and nearly transparent species, the dwarf pygmy goby has a moderately elongated and robust body. Males are slender with nearly straight dorsal and ventral profiles, while the females appear stouter with the dorsal profile slightly curved, the belly protuberant, and the ventral outline strongly arched. The head of the ''P. pygmaea'' is large and blunt. The head and nape are naked. The upper and lower profiles converge and are pointed if viewed from the sides. The head is characteristically broader than deep. The head is very short, broad and rounded. The mouth is very oblique, with a projecting lower jaw and chin. The posterior angle of the maxillary extends beneath the anterior part of the eye, up to the middle of pupil. It possesses two rows of teeth in each jaw. The outer row of teeth inside the upper jaw is larger and more widely spaced, while the inner row is characteristically minute. The first dorsal fin is low, much in advance of the second dorsal fin but not reaching the second dorsal fin when depressed. The pectoral and ventral fins are pointed, with the latter about as long as the former or longer. The anal papilla is very short and rounded in females, but is longer and very slender in males. ''P. pygmaea'' have dark spots, which forms 4 cross-bands, over the sides of its body. The bases of the fins are heavily pigmented, except for the ventral fins. It has 22 to 25 scales in longitudinal series.


Behavior


Feeding

This species feeds on
plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...
.


Reproduction

''P. pygmaea'' is an
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
species.


Miscellaneous

''Pandaka pygmaea'' was once portrayed on a
Philippine The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
ten
centavo The centavo (Spanish and Portuguese 'one hundredth') is a fractional monetary unit that represents one hundredth of a basic monetary unit in many countries around the world. The term comes from Latin ''centum'', ('one hundred'), with the added suf ...
coin where, uniquely for a wildlife portrayal on coinage, it is depicted at life-size. The genus name is derived from the
Filipino Filipino may refer to: * Something from or related to the Philippines ** Filipino language, standardized variety of 'Tagalog', the national language and one of the official languages of the Philippines. ** Filipinos, people who are citizens of th ...
word ''pandak'', meaning "dwarf" or "short of stature".


See also

*
Seven-figure pygmy goby ''Eviota sigillata'', commonly called seven-figure pygmy goby or adorned dwarfgoby, is a species of marine fish in the family Gobiidae. They are widespread throughout the tropical waters of the Indo-West Pacific area, from the Seychelles to the M ...


References


External links

* *
Photograph and Description of Pandaka pygmaea, Herre, 1927, Dwarf pygmy goby, Pandaka pygmaea, NSMT-P 66047, 1.0 cm SL, Fishes of Libong Island, Kahaku.Go.jp
retrieved on July 14, 2007
Pandaka pygmaea, Herre, 1927, Taxonomic Serial No.: 172193, ITIS.gov
retrieved on: July 14, 2007
Chen, Loris. Gobies: Going, Going, Gone? A WebQuest for 6-8th Grade (Science) (lchen@wyckoffschools.org), Ocean Biographic Information System, Census of Marine Life, Rutgers, The State University of New Jersey, 2002-2007. IOBIS.org
retrieved on July 14, 2007

retrieved on July 14, 2007 {{Taxonbar, from=Q244168 Pandaka (fish) Fish of Indonesia Freshwater fish of Indonesia Fish of the Philippines Freshwater fish of the Philippines Fish of Singapore Taxa named by Albert William Herre Fish described in 1927