The lowermost Upper Devonian fossil ''Pallaviciniites'' was for a time the oldest known liverwort until '' Metzgeriothallus'' was recovered from earlier Devonian strata. It had a central axis, and bifurcated at its tips; similar fossils have been found in younger strata through to the Pleistocene. With the exception of its elongated axial conducting (non-vascular) cells, the thallus was a single cell thick. It had a serrated margin. Prior to its discovery, the oldest known liverworts dated to the Lower Carboniferous.

References

Metzgeriales Late Devonian plants Frasnian life Prehistoric plant genera Liverwort genera {{devonian-plant-stub