Paleontology in Colorado
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The location of the state of Colorado Paleontology in Colorado refers to
paleontological Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...
research occurring within or conducted by people from the
U.S. The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
state of
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
. The geologic column of Colorado spans about one third of Earth's history. Fossils can be found almost everywhere in the state but are not evenly distributed among all the ages of the state's rocks. During the early
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
, Colorado was covered by a warm shallow
sea The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...
that would come to be home to creatures like
brachiopods Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, wh ...
,
conodonts Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
,
ostracoderms Ostracoderms () are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes) (may also be polyphyletic if anaspids are closer to cyclostomes) and ...
,
sharks Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorp ...
and
trilobites Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
. This sea withdrew from the state between the Silurian and early Devonian leaving a gap in the local
rock record The geologic record in stratigraphy, paleontology and other natural sciences refers to the entirety of the layers of rock strata. That is, deposits laid down by volcanism or by deposition of sediment derived from weathering detritus (clays, sand ...
. It returned during the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
. Areas of the state not submerged were richly vegetated and inhabited by amphibians that left behind
footprints Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. The ...
that would later fossilize. During the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
, the sea withdrew and
alluvial fan An alluvial fan is an accumulation of sediments that fans outwards from a concentrated source of sediments, such as a narrow canyon emerging from an escarpment. They are characteristic of mountainous terrain in arid to semiarid climates, but a ...
s and sand dunes spread across the state. Many
trace fossils A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils ...
are known from these deposits. The sea returned during the
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
, while exposed areas were a richly vegetated
coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Coa ...
that was home to dinosaurs. Colorado was again submerged by a sea during the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
period that was home to
plesiosaurs The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared i ...
up to 70 feet long. During the early part of the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
era,
rainforests Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
grew in Colorado. Later, another rich flora and fauna would come to be preserved in the Florissant beds, where both
rhinoceroses A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
and uintatheres lived. More recently the state's modern
prairies Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
began to form and the state was home to creatures like bison,
camels A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
, horses, and
mammoths A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, i ...
. Local Native Americans have devised
myths Myth is a folklore genre consisting of Narrative, narratives that play a fundamental role in a society, such as foundational tales or Origin myth, origin myths. Since "myth" is widely used to imply that a story is not Objectivity (philosophy), ...
to explain local fossil bones and dinosaur footprints. By the late 19th century, local fossils had attracted the attention of formally trainer scientists. Major finds include the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
dinosaurs of the Morrison Formation and the Cenozoic plants and mammals of the Florissant beds. The
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
plated dinosaur ''
Stegosaurus armatus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fou ...
'' is the Colorado state fossil. ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been foun ...
'' is also the state dinosaur of Colorado.


Prehistory

No Precambrian fossils are known from Colorado, so the state's fossil record does not begin until the
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
. At the start of the Paleozoic, Colorado was located near the equator. The state was submerged under a warm shallow
sea The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...
. At least part of Colorado was covered by shallow water during the
Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. Th ...
. At the time, Colorado was home to invertebrates like articulated brachiopods,
conodonts Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
,
gastropods The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. The ...
,
ostracods Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typical ...
, pelecypods,
sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through ...
,
trilobites Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
, and
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wo ...
s (known from trace fossils). Contemporary vertebrates included armored
jawless fish Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present ( cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclosto ...
called
ostracoderms Ostracoderms () are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes) (may also be polyphyletic if anaspids are closer to cyclostomes) and ...
. Sometime between the Silurian and early Devonian the sea withdrew from the state. While the sea was gone local sediments were eroded away rather than deposited. During the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
the absent sea returned, although some areas of the state remained dry land. Brachiopods, crinoids, sharks, and trilobites inhabited the sea. A rich variety of plants grew in Colorado's terrestrial environments. Examples include ''
Calamites ''Calamites'' is a genus of extinct arborescent (tree-like) horsetails to which the modern horsetails (genus ''Equisetum'') are closely related. Unlike their herbaceous modern cousins, these plants were medium-sized trees, growing to heights o ...
'',
conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
, and
lycopods Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching s ...
.
Mountain ranges A mountain range or hill range is a series of mountains or hills arranged in a line and connected by high ground. A mountain system or mountain belt is a group of mountain ranges with similarity in form, structure, and alignment that have arise ...
were being raised in the western part of the state by geologic forces. During the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
,
footprints Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. The ...
were laid down in Colorado by early
tetrapods Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids ...
that would later fossilize. That being said, Colorado is not generally a good source of Carboniferous aged fossils. Western Colorado had a series of
alluvial fans An alluvial fan is an accumulation of sediments that fans outwards from a concentrated source of sediments, such as a narrow canyon emerging from an escarpment. They are characteristic of mountainous terrain in arid to semiarid climates, but a ...
during the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
when the
Cutler Group The Cutler Formation or Cutler Group is a rock unit that is exposed across the U.S. states of Arizona, northwest New Mexico, southeast Utah and southwest Colorado. It was laid down in the Early Permian during the Wolfcampian epoch. Descri ...
was being deposited. Preserved in these
sediments Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand a ...
are tracks referred to the
ichnospecies An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
'' Limnopus cutlerensis'', which may have been left by a temnospodyl
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
. During the Permian the sea withdrew once more from the state. In its place were fields of sand dunes. Tracks left by ancient insects and reptiles were preserved in these dune deposits. Also like the Carboniferous, despite the presence of contemporary trace fossils, the fossil record of Permian life in Colorado is relatively poor compared to states like
Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the ...
and
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. Seawater returned to Colorado during the ensuing
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
period, although it left significant areas of the state uncovered. These terrestrial areas included coastal floodplains vegetated by conifers and inhabited by creatures like amphibians and dinosaurs. The
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
also saw the formation of many footprints that would later fossilize. These are preserved in the sediments of the
Chinle Formation The Chinle Formation is an Upper Triassic continental geological formation of fluvial, lacustrine, and palustrine to eolian deposits spread across the U.S. states of Nevada, Utah, northern Arizona, western New Mexico, and western Colorado. In Ne ...
of the northwestern part of the state. A chicken to turkey sized
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
left behind footprints of the ichnogenus '' Agialopous''. These tracks contain large number of the ichnogenus ''
Rhynchosauroides ''Rhynchosauroides'' is an ichnogenus, a form taxon based on footprints. The organism producing the footprints was likely a lepidosaur and may have been a sphenodont, an ancestor of the modern tuatara. The footprint consists of five digits, of w ...
'', which resembles
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
footprints. The Chinle of Colorado also bears the greatest known abundance of the ichnogenus '' Gwynnedichnium''. Both Triassic amphibians and reptiles left behind footprints near what is now the Fall Creek Post Office. During the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
deposition of the sediments now known as the Morrison Formation, both
sauropods Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
and
theropods Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
left behind footprints. Only two large tracksites of fossil footprints are known from the Morrison Formation and both of them are located in Colorado. A tracksite called Rancho del Rio preserves both sauropod and theropod tracks. The Rancho del Rio site is located along the
Colorado River The Colorado River ( es, Río Colorado) is one of the principal rivers (along with the Rio Grande) in the Southwestern United States and northern Mexico. The river drains an expansive, arid drainage basin, watershed that encompasses parts of ...
in central Colorado. The other large tracksite is the Purgatoire Valley tracksite of the eponymous
Purgatoire River The Purgatoire River ( es, Río Purgatorio) is a river in southeastern Colorado, United States. The river is also known locally as the Purgatory River or the Picketwire River. ''Purgatoire'' means Purgatory in French. French trappers named the ri ...
in southeastern Colorado. The Purgatoire Valley tracksite is 400 meters at its widest and contains four track-bearing strata. One of the four track bearing strata bears more than 1,300 individual prints. A series of five parallel trackways left by young sauropods provides important evidence for dinosaur social behavior. The trackways of young sauropods found at the Purgatoire Valley site fill important gaps in the local body fossil record, as the vast majority of sauropods skeletal remains in the Morrison come from grown individuals. Much of Colorado was covered by an expanding sea during the ensuing
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
period. This sea is known as the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea, ...
. Algae, fish, molluscs, and marine reptiles inhabited its waters. The
mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on th ...
''
Platecarpus ''Platecarpus'' ("flat wrist") is an extinct genus of aquatic lizards belonging to the mosasaur family, living around 84–81 million years ago during the middle Santonian to early Campanian, of the Late Cretaceous period. Fossils have been found ...
'' was one such marine reptile. When the
Graneros Shale The Graneros Shale is a geologic formation in the United States identified in the Great Plains as well as New Mexico that dates to the Cenomanian Age of the Cretaceous Period. It is defined as the finely sandy argillaceous or clayey ''near-shore/ ...
was being deposited in Colorado, the
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
'' Thalassomedon'' lived in the state. This was a truly huge plesiosaur that could exceed 45 feet in length. Beyond the shores of this sea were forests and swamps where early flowering plants grew. Later in the Cretaceous, the sea withdrew. Dinosaurs were still present, but the vegetation had undergone significant changes. The forests were now made of broadleafed trees and palms. At this point the local Rocky Mountains began to rise. During the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
cephalopods A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, an ...
with coiled shells and clams were preserved at Monument Creek.
Fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
were present in Coloradan waters and left behind scales that would later fossilize. On land the flora also left behind
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
that would later fossilize. The sediments of the
Benton Formation The Benton Shale (also Benton Formation or Benton Group) is a geologic formation name historically used in Montana, Wyoming, North Dakota, South Dakota, Colorado, Kansas, and Nebraska. In the "mile high" plains in the center of the continent, th ...
preserved both
invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
and
marine reptiles Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the ...
. The
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
'' Ostrea congesta'' was preserved in the Colorado
Niobrara Formation The Niobrara Formation , also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is compose ...
. The Niobrara's vertebrate life included
sharks Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorp ...
, which left behind fossil teeth. Colorado was home to bivalves and straight shelled cephalopods when the Pierre shale was being deposited. The Pierre shale are mound-shaped
bioherm A reef knoll is a land-based landform that comprises an immense pile of calcareous material that accumulated on a previously existing ancient sea floor. At the time of its accumulation it may have had enough structure from organisms such as sponges ...
s up to 15 inches in diameter. A sixty to seventy foot
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
was preserved in what is now
Baca County Baca County is a county located in the U.S. state of Colorado. As of the 2020 census, the population was 3,506. The county seat is Springfield. Located at the southeast corner of Colorado, the county shares state borders with Kansas, New Mexic ...
, which is in the southeastern region of the state. Marine
mollusks Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
were preserved in the
Fox Hills Formation The Fox Hills Formation is a Cretaceous geologic Formation (geology), formation in the northwestern Great Plains of North America. It is present from Alberta on the north to Colorado in the south. Fossil remains of dinosaurs, including tyrannos ...
.
Oysters Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not al ...
and other mollusks were preserved in the sediments now composing the
Dakota Formation The Dakota is a sedimentary geologic unit name of formation and group rank in Midwestern North America. The Dakota units are generally composed of sandstones, mudstones, clays, and shales deposited in the Mid-Cretaceous opening of the Western Int ...
. Aspects of the Coloradan flora were also preserved from this time. The most common plant fossils of the Dakota are the leaves of
deciduous trees In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
. Other contemporary plants included
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s and palms. The uplift of the Rocky Mountains persisted into the early part of the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
era. They were surrounded by
rainforests Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
at this point in prehistory. Areas in the state with lower elevation became the sites of vast lakes. Fish, insects, and leaves would end up entombed in sediments deposited by these lakes. After the start of the Cenozoic, early
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...
s left behind fossils near modern
Golden Golden means made of, or relating to gold. Golden may also refer to: Places United Kingdom *Golden, in the parish of Probus, Cornwall * Golden Cap, Dorset *Golden Square, Soho, London *Golden Valley, a valley on the River Frome in Gloucestershi ...
. The Coloradan flora of the ensuing
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...
left behind plant fossils like ferns, palm leaves, and
petrified wood Petrified wood, also known as petrified tree (from Ancient Greek meaning 'rock' or 'stone'; literally 'wood turned into stone'), is the name given to a special type of ''fossilized wood'', the fossilized remains of terrestrial vegetation. ''P ...
. Animal life of northwestern Colorado during the Eocene included the primitive horse ''
Eohippus ''Eohippus'' is an extinct genus of small equid ungulates. The only species is ''E. angustidens'', which was long considered a species of ''Hyracotherium''. Its remains have been identified in North America and date to the Early Eocene (Ypresian ...
'', early
titanotheres Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to ...
, and uintatheres. A rich flora grew in Colorado during the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
. At least 150 different kinds of plants from this epoch are preserved in what is now the Florissant beds of Colorado. Among the members of this flora were sequoia trees with trunks up to feet in diameter. More than a thousand different kinds of
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
have been documented among the same beds.
Beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s were among the Florissant insects. A diverse mammalian fauna inhabited this ancient forest. Members included animals resembling giant
pig The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus s ...
s,
rhinoceroses A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
, and titanotheres. During the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58
and giant pig-like animals. The state's modern prairies formed during the Quaternary. Colorado was shaken by
volcanic eruptions Several types of volcanic eruptions—during which lava, tephra (ash, lapilli, volcanic bombs and volcanic blocks), and assorted gases are expelled from a volcanic vent or fissure—have been distinguished by volcanologists. These are often ...
. The state's climate gradually cooled as the Cenozoic proceeded. The rainforests gave way to '' sequoia'' forests and
grasslands A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natural ...
.
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
Colorado had a diverse mammal fauna. Among them were ''
Archidiskodon A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...
'', a relative of modern
elephants Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae and ...
.
Bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
,
camels A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
, horses, and
mammoths A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, i ...
also inhabited the state at this time.


History


Indigenous interpretations

Not far from Grand Junction is a tall cottonwood tree called the Ute Council Tree. Close by is an obvious
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
track Track or Tracks may refer to: Routes or imprints * Ancient trackway, any track or trail whose origin is lost in antiquity * Animal track, imprints left on surfaces that an animal walks across * Desire path, a line worn by people taking the shorte ...
site. The Ute had a myth regarding these tracks that justified the tree as a significant meeting place, although the contents of the tale are now lost.
Oglala Lakota The Oglala (pronounced , meaning "to scatter one's own" in Lakota language) are one of the seven subtribes of the Lakota people who, along with the Dakota people, Dakota, make up the Sioux, Očhéthi Šakówiŋ (Seven Council Fires). A majority ...
historian Johnson Holy Rock has described an old story about a Lakota hunting party traveling through northeastern Colorado who were caught during an exceptionally violent thunderstorm while camping. They thought the violence of the storm was due to the Thunder Birds being angry and trying to kill something with their
lightning Lightning is a naturally occurring electrostatic discharge during which two electric charge, electrically charged regions, both in the atmosphere or with one on the land, ground, temporarily neutralize themselves, causing the instantaneous ...
. When morning came, the Lakota hunters went down onto the plain where the storm was most intense. There they found the carcass of an animal with an unusual long nose who was "so strange that they wondered how it managed to eat." This story may be based on the region's abundant fossils. Candidates for the unusual remains found after the storm include
brontotheres Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to ...
,
entelodonts Entelodontidae, the entelodonts, are an extinct family of pig-like artiodactyls (even-toed ungulates) which inhabited the Northern Hemisphere (Asia, Europe, and North America) from the late Eocene to the Middle Miocene epochs, about 38-19 million ...
, or
proboscidean The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From ...
remains. Johnson Holy Rock, who told the story, was inclined to think of the animal as a proboscidean or giant
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inhabit ...
.


Scientific research

Around March 1877 a man named Oramel Lucas discovered sauropod bones in a valley called Garden Park located a few miles north of Canon City. He wrote to
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested ...
and
O. C. Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...
, the famous rival paleontologists of the
bone wars The Bone Wars, also known as the Great Dinosaur Rush, was a period of intense and ruthlessly competitive fossil hunting and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope (of the Acade ...
to alert them about his discovery. Although Marsh never responded, Cope did, and Oramel Lucas and his brother Ira began digging up local fossils and sending them to Cope. These turned out to be the remains of a new species. By August of the same year, Cope had formally named the animal excavated by the Lucas brothers ''
Camarasaurus supremus ''Camarasaurus supremus'' is a species of sauropod dinosaur that lived during the Jurassic period in what is now the western United States. It is the type species of ''Camarasaurus'', which also includes the species ''Camarasaurus grandis'', ...
''. This species may have been the most massive known in the entire Morrison Formation, with estimated putting its body weight at more than 100,000 lbs. Later, a crew working on behalf of O. C. Marsh under
Mudge Mudge is a surname. Notable people with the surname include: Politics * Dirk Mudge (1928–2020), Namibian politician * Geoffrey Mudge, English politician Religion * Enoch Mudge (1776–1850), first Methodist minister reared in New England * ...
and Williston started a quarry nearby. They made several important finds like the new species ''
Allosaurus fragilis ''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch ( Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alludi ...
'' and ''
Diplodocus longus ''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπλ ...
''. Following the initial excavations in the quarry field work stopped until 1883. That year brothers
Marshall Marshall may refer to: Places Australia * Marshall, Victoria, a suburb of Geelong, Victoria Canada * Marshall, Saskatchewan * The Marshall, a mountain in British Columbia Liberia * Marshall, Liberia Marshall Islands * Marshall Islands, an i ...
and Henry Felch reopened excavations there, again on behalf of O. C. Marsh. They worked for five year collecting many dinosaurs already known from the formation, but also the new species ''
Ceratosaurus nasicornis ''Ceratosaurus'' (from Ancient Greek, Greek κέρας/κέρατος, ' meaning "horn" and wikt:σαῦρος, σαῦρος ' meaning "lizard") was a carnivorous Theropoda, theropod dinosaur in the Late Jurassic Period (geology), period (Kim ...
''. In 1890 paleontologist
Charles D. Walcott Charles Doolittle Walcott (March 31, 1850February 9, 1927) was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey.Wonderful Life (book) by Stephen Jay Go ...
found broken pieces of the bony plates embedded in the skin of
Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. Th ...
jawless fish Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present ( cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclosto ...
known as
ostracoderms Ostracoderms () are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes) (may also be polyphyletic if anaspids are closer to cyclostomes) and ...
. These fossils were the oldest known vertebrate remains in the world at the time. The rocks preserving the fossils were gray and reddish sandstone deposited by shallow water. Other fossils found alongside the ostracoderms lived articulated brachiopods,
conodonts Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
,
gastropods The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. The ...
,
ostracods Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typical ...
, pelecypods,
sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through ...
,
trilobites Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...
, and
trace fossils A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils ...
left by
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wo ...
s. After the Felch brothers ended their field work, the so-called Marsh-Felch quarry lay unworked for twelve years. However, in 1900 William Utterback began fieldwork in the area under
John Bell Hatcher John Bell Hatcher (October 11, 1861 – July 3, 1904) was an American paleontologist and fossil hunter known as the "king of collectors" and best known for discovering ''Torosaurus'' and ''Triceratops'', two genera of dinosaurs described by Ot ...
for the
Carnegie Museum of Natural History The Carnegie Museum of Natural History (abbreviated as CMNH) is a natural history museum in the Oakland neighborhood of Pittsburgh, Pennsylvania. It was founded by Pittsburgh-based industrialist Andrew Carnegie in 1896. Housing some 22 millio ...
. In the two ensuing years of field work Utterback found many skeletons of previously known dinosaurs, but also the new genus ''
Haplocanthosaurus ''Haplocanthosaurus'' (meaning "simple spined lizard") is a genus of intermediate sauropod dinosaur. Two species, ''H. delfsi'' and ''H. priscus'', are known from incomplete fossil skeletons. It lived during the late Jurassic period (Kimmeridgia ...
''. This was the smallest known sauropod species of the Morrison Formation. Around 1920 major fossil finds occurred in
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
deposits Colorado shares with
South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota people, Lakota and Dakota peo ...
. Dozens of articulated skeletons and skulls were uncovered. Among them were animals resembling giant
pigs The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus s ...
,
rhinoceroses A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
, and titanotheres. One notable rhinoceros discovery had occurred in
Weld County Weld County is a county located in the U.S. state of Colorado. As of the 2020 census, the population was 328,981. The county seat is Greeley. Weld County comprises the Greeley, CO Metropolitan Statistical Area, which is included in the Denver ...
. In 1925 small duckbilled dinosaurs were discovered in eastern Colorado. Later, in 1955, the
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
uncovered a stone block in south-central Colorado preserving several Eocene ''
Eohippus ''Eohippus'' is an extinct genus of small equid ungulates. The only species is ''E. angustidens'', which was long considered a species of ''Hyracotherium''. Its remains have been identified in North America and date to the Early Eocene (Ypresian ...
'' skeletons. In 1960
Malcolm McKenna Malcolm Carnegie McKenna (1930–2008) was an American paleontologist and author on the subject. Paleontologist McKenna began his paleontology career at the Webb School of California (grades 9-12) in Claremont, California, under noted paleontolo ...
discovered two early
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
turtles Turtles are an order (biology), order of reptiles known as Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) an ...
on behalf of the American Museum of Natural History on South Table Mountain. The following year, curator of the University of Colorado Museum at
Boulder In geology, a boulder (or rarely bowlder) is a rock fragment with size greater than in diameter. Smaller pieces are called cobbles and pebbles. While a boulder may be small enough to move or roll manually, others are extremely massive. In c ...
discovered even more turtles of that age at the same place. In the summer of 1961 a major discovery happened south of
Denver Denver () is a consolidated city and county, the capital, and most populous city of the U.S. state of Colorado. Its population was 715,522 at the 2020 census, a 19.22% increase since 2010. It is the 19th-most populous city in the Unit ...
in Douglas County, at a site known as Lamb Spring. Charles Lamb, the owner of a tract of land used for
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
, was
dredging Dredging is the excavation of material from a water environment. Possible reasons for dredging include improving existing water features; reshaping land and water features to alter drainage, navigability, and commercial use; constructing da ...
out the bottom of a watering hole for the cattle. A great abundance of bones were discovered under the thin layer of mud at the bottom of the watering hole. The owners alerted paleontologists at Denver about the find. The researchers at Denver passed on word of the find to the
Smithsonian Institution The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums and education and research centers, the largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge". Founded ...
of
Washington D.C. ) , image_skyline = , image_caption = Clockwise from top left: the Washington Monument and Lincoln Memorial on the National Mall, United States Capitol, Logan Circle, Jefferson Memorial, White House, Adams Morgan, Na ...
In response, the
National Science Foundation The National Science Foundation (NSF) is an independent agency of the United States government that supports fundamental research and education in all the non-medical fields of science and engineering. Its medical counterpart is the National I ...
funded an expedition coordinated by the Smithsonian Institution to the wateringhole. The lowest bones in the deposit were left by
Columbian mammoths Columbian is the adjective form of Columbia. It may refer to: Buildings * The Columbian Theatre, a music hall in northeastern Kansas * The Columbian (Chicago), a building in Illinois Published works * ''The Columbian'', a daily newspaper pub ...
. Higher in the deposit the excavators uncovered bison,
camels A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
, and horses. By the end of the summer, 13 gigantic cases containing a total of 341 fossil bones were shipped to the
National Museum A national museum is a museum maintained and funded by a national government. In many countries it denotes a museum run by the central government, while other museums are run by regional or local governments. In other countries a much greater numb ...
in Washington D.C. The site is now managed and protected as Lamb Spring Archaeological Preserve. In the spring of 1963, road work in Limon County near the town of Limon uncovered a
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, ...
tooth and
tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share c ...
. In 1965, the
Florissant fossil beds The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones and ...
were proposed as a potential federal preserve. Peter Robinson studied the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
fossil vertebrates of Middle Park during the mid-1960s. These fossils were discovered a short distance northwest of Denver. By 1964 he had discovered the skull of a relatively large camel. Another notable discovery during his research program was the seventh known
microfauna Microfauna (Ancient Greek ''mikros'' "small" + New Latin ''fauna'' "animal") refers to microscopic animals and organisms that exhibit animal-like qualities. Microfauna are represented in the animal kingdom (e.g., nematodes, small arthropods) and t ...
site in the park. More recently, in 1982, the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
plated dinosaur ''
Stegosaurus armatus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fou ...
'' was designated the Colorado state fossil. ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been foun ...
'' was also designated the Colorado state dinosaur that same year.


Protected areas

*
Florissant Fossil Beds National Monument The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones and ...


Paleontologists


Births

Elaine Anderson Elaine Anderson (January 8, 1936 – March 26, 2002) was an American paleontologist. She is best known for her work on vertebrate paleontology. Biography Elaine Anderson was born in Salida, Colorado, on January 8, 1936. She was the only child ...
was born in Salida on January 8,
1936 Events January–February * January 20 – George V of the United Kingdom and the British Dominions and Emperor of India, dies at his Sandringham Estate. The Prince of Wales succeeds to the throne of the United Kingdom as King E ...
. Anderson would come to be known primarily for her book, ''The Pleistocene Mammals of North America'' and her research on Ice Age carnivores. Myra Keen was born in
Colorado Springs Colorado Springs is a home rule municipality in, and the county seat of, El Paso County, Colorado, United States. It is the largest city in El Paso County, with a population of 478,961 at the 2020 United States Census, a 15.02% increase since ...
in
1905 As the second year of the massive Russo-Japanese War begins, more than 100,000 die in the largest world battles of that era, and the war chaos leads to the 1905 Russian Revolution against Nicholas II of Russia (Shostakovich's 11th Symphony i ...
. She would go on to become one of the world's foremost paleomalacologists.


Deaths

Elaine Anderson Elaine Anderson (January 8, 1936 – March 26, 2002) was an American paleontologist. She is best known for her work on vertebrate paleontology. Biography Elaine Anderson was born in Salida, Colorado, on January 8, 1936. She was the only child ...
died in
Denver Denver () is a consolidated city and county, the capital, and most populous city of the U.S. state of Colorado. Its population was 715,522 at the 2020 census, a 19.22% increase since 2010. It is the 19th-most populous city in the Unit ...
on March 26,
2002 File:2002 Events Collage.png, From left, clockwise: The 2002 Winter Olympics are held in Salt Lake City; Queen Elizabeth The Queen Mother and her daughter Princess Margaret, Countess of Snowdon die; East Timor gains East Timor independence, indepe ...
at age 66.
Malcolm McKenna Malcolm Carnegie McKenna (1930–2008) was an American paleontologist and author on the subject. Paleontologist McKenna began his paleontology career at the Webb School of California (grades 9-12) in Claremont, California, under noted paleontolo ...
died in
Boulder In geology, a boulder (or rarely bowlder) is a rock fragment with size greater than in diameter. Smaller pieces are called cobbles and pebbles. While a boulder may be small enough to move or roll manually, others are extremely massive. In c ...
on March 3,
2008 File:2008 Events Collage.png, From left, clockwise: Lehman Brothers went bankrupt following the Subprime mortgage crisis; Cyclone Nargis killed more than 138,000 in Myanmar; A scene from the opening ceremony of the 2008 Summer Olympics in Beijing; ...
. McKenna was best known for publishing a comprehensive classification of mammals.
Charles Repenning Charles Albert Repenning (August 4, 1922 in Oak Park, Illinois – January 5, 2005 in Lakewood, Colorado) was an American paleontologist and zoologist noted for his work on shrews, fossil rodents, modern pinnipeds and their extinct relatives, ...
died in Lakewood on January 5,
2005 File:2005 Events Collage V2.png, From top left, clockwise: Hurricane Katrina in the Gulf of Mexico; the Funeral of Pope John Paul II is held in Vatican City; "Me at the zoo", the first video ever to be uploaded to YouTube; Eris was discovered in ...
. Repenning is known for research into fossil
desmostylians The Desmostylia (from Greek δεσμά ''desma'', "bundle", and στῦλος ''stylos'', "pillar") are an extinct order of aquatic mammals that existed from the early Oligocene (Rupelian) to the late Miocene (Tortonian) (). Desmostylians are ...
and
shrews Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to different ...
.


Natural history museums

*
Denver Museum of Nature and Science The Denver Museum of Nature & Science is a municipal natural history and science museum in Denver, Colorado. It is a resource for informal science education in the Rocky Mountain region. A variety of exhibitions, programs, and activities help mus ...
,
Denver Denver () is a consolidated city and county, the capital, and most populous city of the U.S. state of Colorado. Its population was 715,522 at the 2020 census, a 19.22% increase since 2010. It is the 19th-most populous city in the Unit ...
. * Morrison Natural History Museum,
Morrison Morrison may refer to: People * Morrison (surname), people with the Scottish surname Morrison * Morrison Heady (1829–1915), American poet * Morrison Mann MacBride (1877–1938), Canadian merchant Places in the United States * Morrison, Colorad ...
* Museum of Western Colorado's Dinosaur Journey Museum, Fruita * University of Colorado Museum of Natural History,
Boulder In geology, a boulder (or rarely bowlder) is a rock fragment with size greater than in diameter. Smaller pieces are called cobbles and pebbles. While a boulder may be small enough to move or roll manually, others are extremely massive. In c ...


Notable clubs and associations

* Western Interior Paleontological Society * Denver Gem & Mineral Show


See also

*
Paleontology in Arizona Paleontology in Arizona refers to paleontological research occurring within or conducted by people from the U.S. state of Arizona. The fossil record of Arizona dates to the Precambrian. During the Precambrian, Arizona was home to a shallow sea whi ...
*
Paleontology in Kansas Paleontology in Kansas refers to paleontological research occurring within or conducted by people from the U.S. state of Kansas. Kansas has been the source of some of the most spectacular fossil discoveries in US history. The fossil record of Kansa ...
*
Paleontology in Nebraska The location of the state of Nebraska Paleontology in Nebraska refers to paleontological research occurring within or conducted by people from the U.S. state of Nebraska. Nebraska is world-famous as a source of fossils. During the early Paleozoic ...
*
Paleontology in New Mexico Paleontology in New Mexico refers to paleontological research occurring within or conducted by people from the U.S. state of New Mexico. The fossil record of New Mexico is exceptionally complete and spans almost the entire stratigraphic column. ...
*
Paleontology in Oklahoma Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of ...
*
Paleontology in Utah The location of the state of Utah Paleontology in Utah refers to paleontological research occurring within or conducted by people from the U.S. state of Utah. Utah has a rich fossil record spanning almost all of the geologic column. During the P ...
*
Paleontology in Wyoming Paleontology in Wyoming includes research into the prehistoric life of the U.S. state of Wyoming as well as investigations conducted by Wyomingite researchers and institutions into ancient life occurring elsewhere. The fossil record of the US sta ...


Footnotes


References

* Bell, Christopher J. and Jeanne L. Repenning Forsberg
A Biographical Sketch and Bibliography of Charles A. Repenning
Accessed 12-20-12.

Colorado Department of Personnel and Administration. Accessed 12-20-12.
Directions, Parking & Entrances
Denver Museum of Nature and Science. Accessed 12-20-12. * Everhart, M. J. 2005. ''Oceans of Kansas - A Natural History of the Western Interior Sea''. Indiana University Press, 320 pp. * * Foster, J. (2007). ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World.'' Indiana University Press. 389pp. . * *
Home
Dinosaur Depot Museum. Accessed 12-20-12. * Johnson, Kirk, Dale Springer, Judy Scotchmoor. July 14, 2010.
Colorado, US

The Paleontology Portal
Accessed September 21, 2012. * Lockley, Martin and Hunt, Adrian. '' Dinosaur Tracks of Western North America''. Columbia University Press. 1999. * Mayor, Adrienne. ''Fossil Legends of the First Americans''. Princeton University Press. 2005. .

Morrison Natural History Museum. Accessed 12-20-12.
Plan Your Visit
University of Colorado Museum of Natural History. Accessed 12-20-12. * *
Visit: Dinosaur Journey Museum
Dinosaur Journey Museum. Museum of Western Colorado. Accessed 12-20-12. * Wilford, John Noble

''The New York Times''. March 10, 2008. Accessed 12-20-12.


External links


Geologic units in Colorado

Paleoportal: Colorado
{{Colorado
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
Natural history of Colorado Science and technology in Colorado