Paleolithic Age
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The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''
palaios ''PALAIOS'' is a bimonthly academic journal dedicated to the study of the impact of life on Earth history, combining the fields of palaeontology and sedimentology. It has been published by the Society for Sedimentary Geology since its inception i ...
'', "old" and λίθος '' lithos'', "stone"), is a period in human
prehistory Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use ...
that is distinguished by the original development of stone tools, and which represents almost the entire period of human
prehistoric technology Prehistoric technology is technology that predates recorded history. History is the study of the past using written records. Anything prior to the first written accounts of history is prehistoric, including earlier technologies. About 2.5 million ...
. It extends from the earliest known use of stone tools by
hominins The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The t ...
,  3.3 million years ago, to the end of the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
,  11,650 cal BP. The Paleolithic Age in Europe preceded the
Mesolithic Age The Mesolithic (Greek: μέσος, ''mesos'' 'middle' + λίθος, ''lithos'' 'stone') or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic is often used synonymously ...
, although the date of the transition varies geographically by several thousand years. During the Paleolithic Age, hominins grouped together in small societies such as bands and subsisted by gathering plants, fishing, and hunting or scavenging wild animals. The Paleolithic Age is characterized by the use of
knapped Knapping is the shaping of flint, chert, obsidian, or other conchoidal fracturing stone through the process of lithic reduction to manufacture stone tools, strikers for flintlock firearms, or to produce flat-faced stones for building or facing w ...
stone tools, although at the time humans also used wood and bone tools. Other organic commodities were adapted for use as tools, including
leather Leather is a strong, flexible and durable material obtained from the tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffalo, pigs and hog ...
and vegetable
fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorpora ...
s; however, due to rapid decomposition, these have not survived to any great degree. About 50,000 years ago, a marked increase in the diversity of artifacts occurred. In Africa, bone artifacts and the first
art Art is a diverse range of human activity, and resulting product, that involves creative or imaginative talent expressive of technical proficiency, beauty, emotional power, or conceptual ideas. There is no generally agreed definition of wha ...
appear in the archaeological record. The first evidence of human
fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques inclu ...
is also noted, from artifacts in places such as Blombos cave in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. Archaeologists classify artifacts of the last 50,000 years into many different categories, such as
projectile point In North American archaeological terminology, a projectile point is an object that was hafted to a weapon that was capable of being thrown or projected, such as a javelin, dart, or arrow. They are thus different from weapons presumed to have ...
s, engraving tools, Sharp knife blades, and drilling and piercing tools. Humankind gradually evolved from early members of the genus ''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely relat ...
''—such as ''Homo habilis'', who used simple stone tools—into
anatomically modern humans Early modern human (EMH) or anatomically modern human (AMH) are terms used to distinguish '' Homo sapiens'' (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans from exti ...
as well as behaviourally modern humans by the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
. During the end of the Paleolithic Age, specifically the Middle or Upper Paleolithic Age, humans began to produce the earliest works of art and to engage in
religious Religion is usually defined as a social- cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatur ...
or spiritual behavior such as burial and
ritual A ritual is a sequence of activities involving gestures, words, actions, or objects, performed according to a set sequence. Rituals may be prescribed by the traditions of a community, including a religious community. Rituals are characterized ...
. Conditions during the Paleolithic Age went through a set of glacial and
interglacial period An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene in ...
s in which the
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologi ...
periodically fluctuated between warm and cool temperatures. Archaeological and genetic data suggest that the source populations of Paleolithic humans survived in sparsely-wooded areas and dispersed through areas of high
primary productivity In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through c ...
while avoiding dense forest-cover. By  BP, the first humans set foot in Australia. By  BP, humans lived at 61°N latitude in Europe. By  BP, Japan was reached, and by  BP humans were present in
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
, above the Arctic Circle. By the end of the Upper Paleolithic Age humans had crossed Beringia and expanded throughout the Americas.


Etymology

The term "
Palaeolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''palaios'', "old" and λίθος '' lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone to ...
" was coined by archaeologist John Lubbock in 1865. It derives from Greek: παλαιός, ''palaios'', "old"; and λίθος, ''lithos'', "stone", meaning "old age of the stone" or "Old Stone Age".


Paleogeography and climate

The Paleolithic coincides almost exactly with the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
epoch of geologic time, which lasted from 2.6 million years ago to about 12,000 years ago. This epoch experienced important geographic and climatic changes that affected human societies. During the preceding
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58drift from possibly as far as from their present locations to positions only from their current location. South America became linked to North America through the Isthmus of Panama, bringing a nearly complete end to South America's distinctive
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
fauna. The formation of the isthmus had major consequences on global temperatures, because warm equatorial ocean currents were cut off, and the cold Arctic and Antarctic waters lowered temperatures in the now-isolated Atlantic Ocean. Most of
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
formed during the Pliocene to connect the continents of North and South America, allowing fauna from these continents to leave their native habitats and colonize new areas. Africa's collision with Asia created the Mediterranean, cutting off the remnants of the
Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
. During the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
, the modern
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions In geography, regions, otherwise referred to as zones, lands or territories, are areas t ...
s were essentially at their present positions; the
tectonic plate Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large te ...
s on which they sit have probably moved at most from each other since the beginning of the period. Climates during the Pliocene became cooler and drier, and seasonal, similar to modern climates.
Ice sheet In glaciology, an ice sheet, also known as a continental glacier, is a mass of glacial ice that covers surrounding terrain and is greater than . The only current ice sheets are in Antarctica and Greenland; during the Last Glacial Period at La ...
s grew on
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
. The formation of an Arctic ice cap around 3 million years ago is signaled by an abrupt shift in
oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as ...
isotope Isotopes are two or more types of atoms that have the same atomic number (number of protons in their nuclei) and position in the periodic table (and hence belong to the same chemical element), and that differ in nucleon numbers (mass numb ...
ratios and ice-rafted cobbles in the North Atlantic and North
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
beds. Mid-latitude
glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate be ...
probably began before the end of the epoch. The global cooling that occurred during the Pliocene may have spurred on the disappearance of forests and the spread of
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s and
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s. The
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
climate was characterized by repeated glacial cycles during which
continental glacier In glaciology, an ice sheet, also known as a continental glacier, is a mass of glacial ice that covers surrounding terrain and is greater than . The only current ice sheets are in Antarctica and Greenland; during the Last Glacial Period at La ...
s pushed to the 40th parallel in some places. Four major glacial events have been identified, as well as many minor intervening events. A major event is a general glacial excursion, termed a "glacial". Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials". Each glacial advance tied up huge volumes of water in continental ice sheets deep, resulting in temporary sea level drops of or more over the entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions. The effects of glaciation were global.
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
was ice-bound throughout the Pleistocene and the preceding Pliocene. The
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
were covered in the south by the
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
n ice cap. There were glaciers in New Zealand and
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
. The now decaying glaciers of Mount Kenya, Mount Kilimanjaro, and the Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
and to the west in the Atlas mountains. In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the
Laurentide The Laurentide Ice Sheet was a massive sheet of ice that covered millions of square miles, including most of Canada and a large portion of the Northern United States, multiple times during the Quaternary glacial epochs, from 2.58 million years a ...
covered the east. The Fenno-Scandian ice sheet covered northern Europe, including Great Britain; the Alpine ice sheet covered the Alps. Scattered domes stretched across
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
and the Arctic shelf. The northern seas were frozen. During the late Upper Paleolithic (Latest Pleistocene)  BP, the Beringia land bridge between Asia and North America was blocked by ice, which may have prevented early Paleo-Indians such as the
Clovis culture The Clovis culture is a prehistoric Paleoamerican culture, named for distinct stone and bone tools found in close association with Pleistocene fauna, particularly two mammoths, at Blackwater Locality No. 1 near Clovis, New Mexico, in 1936 a ...
from directly crossing Beringia to reach the Americas. According to Mark Lynas (through collected data), the Pleistocene's overall climate could be characterized as a continuous
El Niño El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date ...
with
trade winds The trade winds or easterlies are the permanent east-to-west prevailing winds that flow in the Earth's equatorial region. The trade winds blow mainly from the northeast in the Northern Hemisphere and from the southeast in the Southern Hemisp ...
in the south
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
weakening or heading east, warm air rising near
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
, warm water spreading from the west Pacific and the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by t ...
to the east Pacific, and other El Niño markers. The Paleolithic is often held to finish at the end of the ice age (the end of the Pleistocene epoch), and Earth's climate became warmer. This may have caused or contributed to the extinction of the
Pleistocene megafauna Pleistocene megafauna is the set of large animals that lived on Earth during the Pleistocene epoch. Pleistocene megafauna became extinct during the Quaternary extinction event resulting in substantial changes to ecosystems globally. The role of ...
, although it is also possible that the late
Pleistocene extinctions The Quaternary period (from 2.588 ± 0.005 million years ago to the present) has seen the extinctions of numerous predominantly megafaunal species, which have resulted in a collapse in faunal density and diversity and the extinction of key ecolog ...
were (at least in part) caused by other factors such as disease and overhunting by humans. New research suggests that the extinction of the
woolly mammoth The woolly mammoth (''Mammuthus primigenius'') is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus s ...
may have been caused by the combined effect of climatic change and human hunting. Scientists suggest that climate change during the end of the Pleistocene caused the mammoths' habitat to shrink in size, resulting in a drop in population. The small populations were then hunted out by Paleolithic humans. The global warming that occurred during the end of the Pleistocene and the beginning of the
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
may have made it easier for humans to reach mammoth habitats that were previously frozen and inaccessible. Small populations of woolly mammoths survived on isolated Arctic islands, Saint Paul Island and
Wrangel Island Wrangel Island ( rus, О́стров Вра́нгеля, r=Ostrov Vrangelya, p=ˈostrəf ˈvrangʲɪlʲə; ckt, Умӄиԓир, translit=Umqiḷir) is an island of the Chukotka Autonomous Okrug, Russia. It is the 91st largest island in the w ...
, until  BP and  BP respectively. The Wrangel Island population became extinct around the same time the island was settled by prehistoric humans. There is no evidence of prehistoric human presence on Saint Paul island (though early human settlements dating as far back as 6500 BP were found on the nearby
Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a chain of 14 large v ...
).


Human way of life

Nearly all of our knowledge of Paleolithic human culture and way of life comes from
archaeology Archaeology or archeology is the scientific study of human activity through the recovery and analysis of material culture. The archaeological record consists of artifacts, architecture, biofacts or ecofacts, sites, and cultural landsca ...
and
ethnographic Ethnography (from Greek ''ethnos'' "folk, people, nation" and ''grapho'' "I write") is a branch of anthropology and the systematic study of individual cultures. Ethnography explores cultural phenomena from the point of view of the subject ...
comparisons to modern hunter-gatherer cultures such as the !Kung San who live similarly to their Paleolithic predecessors. The economy of a typical Paleolithic society was a hunter-gatherer economy. Humans hunted wild animals for meat and gathered food, firewood, and materials for their tools, clothes, or shelters. Human population density was very low, around only . This was most likely due to low body fat, infanticide, high levels of physical activity among women, late weaning of infants, and a
nomad A nomad is a member of a community without fixed habitation who regularly moves to and from the same areas. Such groups include hunter-gatherers, pastoral nomads (owning livestock), tinkers and trader nomads. In the twentieth century, the po ...
ic lifestyle. Like contemporary hunter-gatherers, Paleolithic humans enjoyed an abundance of leisure time unparalleled in both
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several p ...
farming societies and modern industrial societies. At the end of the Paleolithic, specifically the Middle or Upper Paleolithic, humans began to produce works of art such as cave paintings, rock art and
jewellery Jewellery ( UK) or jewelry (U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a wester ...
and began to engage in religious behavior such as burials and rituals.


Distribution

At the beginning of the Paleolithic, hominins were found primarily in eastern Africa, east of the
Great Rift Valley The Great Rift Valley is a series of contiguous geographic trenches, approximately in total length, that runs from Lebanon in Asia to Mozambique in Southeast Africa. While the name continues in some usages, it is rarely used in geology as it ...
. Most known hominin fossils dating earlier than one million years before present are found in this area, particularly in
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
, and
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
. By  BP, groups of hominins began leaving Africa and settling southern Europe and Asia. Southern Caucasus was occupied by  BP, and northern China was reached by  BP. By the end of the Lower Paleolithic, members of the hominin family were living in what is now China, western Indonesia, and, in Europe, around the Mediterranean and as far north as England, France, southern Germany, and Bulgaria. Their further northward expansion may have been limited by the lack of control of fire: studies of cave settlements in Europe indicate no regular use of fire prior to  BP. East Asian fossils from this period are typically placed in the genus '' Homo erectus''. Very little fossil evidence is available at known Lower Paleolithic sites in Europe, but it is believed that hominins who inhabited these sites were likewise ''Homo erectus''. There is no evidence of hominins in America, Australia, or almost anywhere in Oceania during this time period. Fates of these early colonists, and their relationships to modern humans, are still subject to debate. According to current archaeological and genetic models, there were at least two notable expansion events subsequent to peopling of Eurasia  BP. Around 500,000 BP a group of early humans, frequently called ''
Homo heidelbergensis ''Homo heidelbergensis'' (also ''H. sapiens heidelbergensis''), sometimes called Heidelbergs, is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of '' H. erectus'' i ...
'', came to Europe from Africa and eventually evolved into ''Homo neanderthalensis'' (
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While the ...
s). In the Middle Paleolithic, Neanderthals were present in the region now occupied by Poland. Both ''Homo erectus'' and ''Homo neanderthalensis'' became extinct by the end of the Paleolithic. Descended from ''Homo sapiens'', the anatomically modern ''
Homo sapiens sapiens Human taxonomy is the classification of the human species (systematic name ''Homo sapiens'', Latin: "wise man") within zoological taxonomy. The systematic genus, '' Homo'', is designed to include both anatomically modern humans and extinct ...
'' emerged in eastern Africa  BP, left Africa around 50,000 BP, and expanded throughout the planet. Multiple hominid groups coexisted for some time in certain locations. ''Homo neanderthalensis'' were still found in parts of Eurasia  BP years, and engaged in an unknown degree of interbreeding with ''Homo sapiens sapiens''. DNA studies also suggest an unknown degree of interbreeding between ''Homo sapiens sapiens'' and ''
Homo sapiens denisova The Denisovans or Denisova hominins ) are an extinct species or subspecies of archaic human that ranged across Asia during the Lower and Middle Paleolithic. Denisovans are known from few physical remains and consequently, most of what is known ...
''. Hominin fossils not belonging either to ''Homo neanderthalensis'' or to ''Homo sapiens'' species, found in the Altai Mountains and Indonesia, were radiocarbon dated to  BP and  BP respectively. For the duration of the Paleolithic, human populations remained low, especially outside the equatorial region. The entire population of Europe between 16,000 and 11,000 BP likely averaged some 30,000 individuals, and between 40,000 and 16,000 BP, it was even lower at 4,000–6,000 individuals. However, remains of thousands of butchered animals and tools made by Palaeolithic humans were found in Lapa do Picareiro ( pt), a cave in
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
, dating back between 41,000 and 38,000 years ago.


Technology


Tools

Paleolithic humans made tools of stone, bone (primarily deer), and wood. The early paleolithic hominins, ''
Australopithecus ''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Austral ...
'', were the first users of stone tools. Excavations in
Gona, Ethiopia Gona is an archaeological site in the Afar Triangle of Ethiopia located in the Ethiopian Lowlands. The site, near the Middle Awash and Hadar regions, is primarily known for paleoanthropological study, including excavations of Late Miocene and E ...
have produced thousands of artifacts, and through radioisotopic dating and
magnetostratigraphy Magnetostratigraphy is a geophysical correlation technique used to date sedimentary and volcanic sequences. The method works by collecting oriented samples at measured intervals throughout the section. The samples are analyzed to determine their '' ...
, the sites can be firmly dated to 2.6 million years ago. Evidence shows these early hominins intentionally selected raw stone with good flaking qualities and chose appropriate sized stones for their needs to produce sharp-edged tools for cutting. The earliest Paleolithic stone tool industry, the
Oldowan The Oldowan (or Mode I) was a widespread stone tool archaeological industry (style) in prehistory. These early tools were simple, usually made with one or a few flakes chipped off with another stone. Oldowan tools were used during the Lower ...
, began around 2.6 million years ago. It produced tools such as choppers, burins, and
stitching awl A stitching awl is a tool with which holes can be punctured in a variety of materials, or existing holes can be enlarged. It is also used for sewing heavy materials, such as leather or canvas. It is a thin, tapered metal shaft, coming to a shar ...
s. It was completely replaced around 250,000 years ago by the more complex
Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...
industry, which was first conceived by ''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unresol ...
'' around 1.8–1.65 million years ago. The Acheulean implements completely vanish from the archaeological record around 100,000 years ago and were replaced by more complex Middle Paleolithic tool kits such as the
Mousterian The Mousterian (or Mode III) is an archaeological industry of stone tools, associated primarily with the Neanderthals in Europe, and to the earliest anatomically modern humans in North Africa and West Asia. The Mousterian largely defines the l ...
and the
Aterian The Aterian is a Middle Stone Age (or Middle Palaeolithic) stone tool industry centered in North Africa, from Mauritania to Egypt, but also possibly found in Oman and the Thar Desert. The earliest Aterian dates to c. 150,000 years ago, at the sit ...
industries. Lower Paleolithic humans used a variety of stone tools, including
hand axe A hand axe (or handaxe or Acheulean hand axe) is a prehistoric stone tool with two faces that is the longest-used tool in human history, yet there is no academic consensus on what they were used for. It is made from stone, usually flint or ch ...
s and
choppers Chopper may refer to: Arts and entertainment * ''Chopper'' (comics), a horror comic book mini-series * ''Chopper'' (film), a 2000 Australian film by and about Mark "Chopper" Read * Chopper (''Judge Dredd''), a character in British comics anth ...
. Although they appear to have used hand axes often, there is disagreement about their use. Interpretations range from cutting and chopping tools, to digging implements, to flaking cores, to the use in traps, and as a purely ritual significance, perhaps in courting behavior. William H. Calvin has suggested that some hand axes could have served as "killer
Frisbees A frisbee (pronounced ), also called a flying disc or simply a disc, is a gliding toy or sporting item that is generally made of injection-molded plastic and roughly in diameter with a pronounced lip. It is used recreationally and competitive ...
" meant to be thrown at a herd of animals at a waterhole so as to stun one of them. There are no indications of
hafting Hafting is a process by which an artifact, often bone, stone, or metal is attached to a ''haft'' (handle or strap). This makes the artifact more useful by allowing it to be shot (arrow), thrown by hand (spear), or used with more effective levera ...
, and some artifacts are far too large for that. Thus, a thrown hand axe would not usually have penetrated deeply enough to cause very serious injuries. Nevertheless, it could have been an effective weapon for defense against predators. Choppers and scrapers were likely used for skinning and butchering scavenged animals and sharp-ended sticks were often obtained for digging up edible roots. Presumably, early humans used wooden spears as early as 5 million years ago to hunt small animals, much as their relatives,
chimpanzees The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
, have been observed to do in
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
, Africa. Lower Paleolithic humans constructed shelters, such as the possible wood hut at Terra Amata.


Fire use

Fire was used by the Lower Paleolithic hominins '' Homo erectus'' and ''
Homo ergaster ''Homo ergaster'' is an extinct species or subspecies of archaic humans who lived in Africa in the Early Pleistocene. Whether ''H. ergaster'' constitutes a species of its own or should be subsumed into '' H. erectus'' is an ongoing and unresol ...
'' as early as 300,000 to 1.5 million years ago and possibly even earlier by the early Lower Paleolithic (Oldowan) hominin '' Homo habilis'' or by robust ''
Australopithecine Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', '' Paranthropus'', and ''Kenyanthropus''), and it typically includ ...
s'' such as '' Paranthropus''. However, the use of fire only became common in the societies of the following Middle Stone Age and Middle Paleolithic. Use of fire reduced mortality rates and provided protection against predators. Early hominins may have begun to cook their food as early as the Lower Paleolithic ( million years ago) or at the latest in the early Middle Paleolithic ( years ago). Some scientists have hypothesized that hominins began cooking food to defrost frozen meat, which would help ensure their survival in cold regions. Archaeologists cite morphological shifts in cranial anatomy as evidence for emergence of cooking and food processing technologies. These morphological changes include decreases in molar and jaw size, thinner tooth enamel, and decrease in gut volume During much of the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
epoch, our ancestors relied on simple food processing techniques such as roasting The
Upper Palaeolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
saw the emergence of boiling, an advance in food processing technology which rendered plant foods more digestible, decreased their toxicity, and maximised their nutritional value Thermally altered rock (heated stones) are easily identifiable in the archaeological record. Stone-boiling and pit-baking were common techniques which involved heating large pebbles then transferring the hot stones into a perishable container to heat the water This technology is typified in the
Middle Palaeolithic The Middle Paleolithic (or Middle Palaeolithic) is the second subdivision of the Paleolithic or Old Stone Age as it is understood in Europe, Africa and Asia. The term Middle Stone Age is used as an equivalent or a synonym for the Middle Pale ...
example of the Abri Pataud hearths


Raft

The Lower Paleolithic ''Homo erectus'' possibly invented
raft A raft is any flat structure for support or transportation over water. It is usually of basic design, characterized by the absence of a hull. Rafts are usually kept afloat by using any combination of buoyant materials such as wood, sealed barrels ...
s ( BP) to travel over large bodies of water, which may have allowed a group of ''Homo erectus'' to reach the island of Flores and evolve into the small hominin ''
Homo floresiensis ''Homo floresiensis'' also known as "Flores Man"; nicknamed "Hobbit") is an extinct species of small archaic human that inhabited the island of Flores, Indonesia, until the arrival of modern humans about 50,000 years ago. The remains of an in ...
''. However, this hypothesis is disputed within the anthropological community. The possible use of rafts during the Lower Paleolithic may indicate that Lower Paleolithic hominins such as ''Homo erectus'' were more advanced than previously believed, and may have even spoken an early form of modern language. Supplementary evidence from Neanderthal and modern human sites located around the Mediterranean Sea, such as Coa de sa Multa ( BP), has also indicated that both Middle and Upper Paleolithic humans used rafts to travel over large bodies of water (i.e. the Mediterranean Sea) for the purpose of colonizing other bodies of land.


Advanced tools

By around 200,000 BP, Middle Paleolithic stone tool manufacturing spawned a tool making technique known as the prepared-core technique, that was more elaborate than previous
Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...
techniques. This technique increased efficiency by allowing the creation of more controlled and consistent flakes. It allowed Middle Paleolithic humans to create stone tipped
spear A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head. The head may be simply the sharpened end of the shaft itself, as is the case with fire hardened spears, or it may be made of a more durable material fasten ...
s, which were the earliest composite tools, by hafting sharp, pointy stone flakes onto wooden shafts. In addition to improving tool making methods, the Middle Paleolithic also saw an improvement of the tools themselves that allowed access to a wider variety and amount of food sources. For example,
microlith A microlith is a small stone tool usually made of flint or chert and typically a centimetre or so in length and half a centimetre wide. They were made by humans from around 35,000 to 3,000 years ago, across Europe, Africa, Asia and Australia. Th ...
s or small stone tools or points were invented around 70,000–65,000 BP and were essential to the invention of bows and
spear thrower A spear-thrower, spear-throwing lever or ''atlatl'' (pronounced or ; Nahuatl ''ahtlatl'' ) is a tool that uses leverage to achieve greater velocity in dart or javelin-throwing, and includes a bearing surface which allows the user to store ene ...
s in the following Upper Paleolithic.
Harpoon A harpoon is a long spear-like instrument and tool used in fishing, whaling, sealing, and other marine hunting to catch and injure large fish or marine mammals such as seals and whales. It accomplishes this task by impaling the target animal ...
s were invented and used for the first time during the late Middle Paleolithic ( BP); the invention of these devices brought fish into the human diets, which provided a hedge against starvation and a more abundant food supply."Human Evolution," Microsoft Encarta Online Encyclopedia 2007
Contributed by Richard B. Potts, B.A., Ph.D.
Thanks to their technology and their advanced social structures, Paleolithic groups such as the Neanderthals—who had a Middle Paleolithic level of technology—appear to have hunted large game just as well as Upper Paleolithic modern humans. and the Neanderthals in particular may have likewise hunted with projectile weapons. Nonetheless, Neanderthal use of projectile weapons in hunting occurred very rarely (or perhaps never) and the Neanderthals hunted large game animals mostly by
ambush An ambush is a long-established military tactic in which a combatant uses an advantage of concealment or the element of surprise to attack unsuspecting enemy combatants from concealed positions, such as among dense underbrush or behind mo ...
ing them and attacking them with mêlée weapons such as thrusting spears rather than attacking them from a distance with projectile weapons.


Other inventions

During the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
, further inventions were made, such as the
net Net or net may refer to: Mathematics and physics * Net (mathematics), a filter-like topological generalization of a sequence * Net, a linear system of divisors of dimension 2 * Net (polyhedron), an arrangement of polygons that can be folded up ...
( or  BP)
bolas Bolas or bolases (singular bola; from Spanish and Portuguese ''bola'', "ball", also known as a ''boleadora'' or ''boleadeira'') is a type of throwing weapon made of weights on the ends of interconnected cords, used to capture animals by entan ...
, the
spear thrower A spear-thrower, spear-throwing lever or ''atlatl'' (pronounced or ; Nahuatl ''ahtlatl'' ) is a tool that uses leverage to achieve greater velocity in dart or javelin-throwing, and includes a bearing surface which allows the user to store ene ...
( BP), the bow and arrow ( or  BP) and the oldest example of ceramic art, the
Venus of Dolní Věstonice The Venus of Dolní Věstonice ( cs, Věstonická venuše) is a Venus figurine, a ceramic statuette of a nude female figure dated to 29,000–25,000 BCE (Gravettian industry). It was found at the Paleolithic site Dolní Věstonice in the Moravian ...
( BP). Kilu Cave at Buku island,
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
, demonstrates navigation of some 60 km of open ocean at 30,000 BCcal. Early dogs were domesticated sometime between 30,000 and 14,000 BP, presumably to aid in hunting. Lloyd, J & Mitchinson, J: "
The Book of General Ignorance ''The Book of General Ignorance'' is the first in a series of books based on the final round in the intellectual British panel game '' QI'', written by series-creator John Lloyd and head-researcher John Mitchinson,
". Faber & Faber, 2006.
However, the earliest instances of successful domestication of dogs may be much more ancient than this. Evidence from canine DNA collected by Robert K. Wayne suggests that dogs may have been first domesticated in the late Middle Paleolithic around 100,000 BP or perhaps even earlier. Archaeological evidence from the
Dordogne Dordogne ( , or ; ; oc, Dordonha ) is a large rural department in Southwestern France, with its prefecture in Périgueux. Located in the Nouvelle-Aquitaine region roughly half-way between the Loire Valley and the Pyrenees, it is name ...
region of France demonstrates that members of the European early
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
culture known as the Aurignacian used calendars ( BP). This was a lunar calendar that was used to document the phases of the moon. Genuine solar calendars did not appear until the Neolithic. Upper Paleolithic cultures were probably able to time the migration of game animals such as wild horses and deer."Stone Age," Microsoft Encarta Online Encyclopedia 2007
Contributed by Kathy Schick, B.A., M.A., Ph.D. and Nicholas Toth, B.A., M.A., Ph.D.
This ability allowed humans to become efficient hunters and to exploit a wide variety of game animals. Recent research indicates that the Neanderthals timed their hunts and the migrations of game animals long before the beginning of the Upper Paleolithic.


Social organization

The social organization of the earliest Paleolithic ( Lower Paleolithic) societies remains largely unknown to scientists, though Lower Paleolithic hominins such as ''Homo habilis'' and ''Homo erectus'' are likely to have had more complex social structures than chimpanzee societies. Late Oldowan/Early Acheulean humans such as ''Homo ergaster''/''Homo erectus'' may have been the first people to invent central campsites or home bases and incorporate them into their foraging and hunting strategies like contemporary hunter-gatherers, possibly as early as 1.7 million years ago; however, the earliest solid evidence for the existence of home bases or central campsites (hearths and shelters) among humans only dates back to 500,000 years ago. Similarly, scientists disagree whether Lower Paleolithic humans were largely monogamous or polygynous. In particular, the Provisional model suggests that
bipedalism Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
arose in pre-Paleolithic
australopithecine Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', '' Paranthropus'', and ''Kenyanthropus''), and it typically includ ...
societies as an adaptation to monogamous lifestyles; however, other researchers note that
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
is more pronounced in Lower Paleolithic humans such as ''Homo erectus'' than in modern humans, who are less polygynous than other primates, which suggests that Lower Paleolithic humans had a largely polygynous lifestyle, because species that have the most pronounced sexual dimorphism tend more likely to be polygynous. Human societies from the Paleolithic to the early Neolithic farming tribes lived without states and organized governments. For most of the Lower Paleolithic, human societies were possibly more hierarchical than their Middle and Upper Paleolithic descendants, and probably were not grouped into bands,Christopher Boehm (1999
"Hierarchy in the Forest: The Evolution of Egalitarian Behavior" pp. 198–208
Harvard University Press
though during the end of the Lower Paleolithic, the latest populations of the hominin ''Homo erectus'' may have begun living in small-scale (possibly egalitarian) bands similar to both Middle and Upper Paleolithic societies and modern hunter-gatherers. Middle Paleolithic societies, unlike Lower Paleolithic and early Neolithic ones, consisted of bands that ranged from 20–30 or 25–100 members and were usually nomadic. These bands were formed by several families. Bands sometimes joined together into larger "macrobands" for activities such as acquiring mates and celebrations or where resources were abundant. By the end of the Paleolithic era ( BP), people began to settle down into permanent locations, and began to rely on agriculture for sustenance in many locations. Much evidence exists that humans took part in long-distance trade between bands for rare commodities (such as ochre, which was often used for religious purposes such as ritual

/ref>) and raw materials, as early as 120,000 years ago in Middle Paleolithic. Inter-band trade may have appeared during the Middle Paleolithic because trade between bands would have helped ensure their survival by allowing them to exchange resources and commodities such as raw materials during times of relative scarcity (i.e. famine, drought). Like in modern hunter-gatherer societies, individuals in Paleolithic societies may have been subordinate to the band as a whole.pp. 9–13p. 70
/ref> Both Neanderthals and modern humans took care of the elderly members of their societies during the Middle and Upper Paleolithic. Some sources claim that most Middle and Upper Paleolithic societies were possibly fundamentally egalitarianism, egalitarianChristopher Boehm (1999
"Hierarchy in the Forest: The Evolution of Egalitarian Behavior" p. 198
Harvard University Press
and may have rarely or never engaged in organized violence between groups (i.e. war).pp. 420-22
/ref>p. 123
/ref> Some Upper Paleolithic societies in resource-rich environments (such as societies in
Sungir Sungir (, sometimes spelled Sunghir) is an Upper Paleolithic archaeological site in Russia and one of the earliest records of modern ''Homo sapiens'' in Eurasia. It is situated about two hundred kilometres east of Moscow, on the outskirts of Vlad ...
, in what is now Russia) may have had more complex and hierarchical organization (such as
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
s with a pronounced hierarchy and a somewhat formal division of labor) and may have engaged in
endemic warfare __NOTOC__ Ritual warfare (sometimes called endemic warfare) is a state of continual or frequent warfare, such as is found in some tribal societies (but is not limited to tribal societies). Description Ritual fighting (or ritual battle or ritual ...
. Some argue that there was no formal leadership during the Middle and Upper Paleolithic. Like contemporary egalitarian hunter-gatherers such as the
Mbuti The Mbuti people, or Bambuti, are one of several indigenous pygmy groups in the Congo region of Africa. Their languages are Central Sudanic languages and Bantu languages. Subgroups Bambuti are pygmy hunter-gatherers, and are one of the old ...
pygmies, societies may have made decisions by communal consensus decision making rather than by appointing permanent rulers such as chiefs and
monarch A monarch is a head of stateWebster's II New College DictionarMonarch Houghton Mifflin. Boston. 2001. p. 707. for life or until abdication, and therefore the head of state of a monarchy. A monarch may exercise the highest authority and power i ...
s. Nor was there a formal division of labor during the Paleolithic. Each member of the group was skilled at all tasks essential to survival, regardless of individual abilities. Theories to explain the apparent egalitarianism have arisen, notably the Marxist concept of primitive communism. Christopher Boehm (1999) has hypothesized that egalitarianism may have evolved in Paleolithic societies because of a need to distribute resources such as food and meat equally to avoid famine and ensure a stable food supply.Christopher Boehm (1999
"Hierarchy in the Forest: The Evolution of Egalitarian Behavior" p. 192
Harvard university press
Raymond C. Kelly speculates that the relative peacefulness of Middle and Upper Paleolithic societies resulted from a low population density, cooperative relationships between groups such as reciprocal exchange of commodities and collaboration on hunting expeditions, and because the invention of projectile weapons such as throwing spears provided less incentive for war, because they increased the damage done to the attacker and decreased the relative amount of territory attackers could gain. However, other sources claim that most Paleolithic groups may have been larger, more complex, sedentary and warlike than most contemporary hunter-gatherer societies, due to occupying more resource-abundant areas than most modern hunter-gatherers who have been pushed into more marginal habitats by agricultural societies. Anthropologists have typically assumed that in Paleolithic societies, women were responsible for gathering wild plants and firewood, and men were responsible for hunting and scavenging dead animals. However, analogies to existent hunter-gatherer societies such as the
Hadza people The Hadza, or Hadzabe (''Wahadzabe'' in Swahili), are a Tanzanian indigenous ethnic group mostly based in southwest Karatu District of Arusha Region. They live around Lake Eyasi in the central Rift Valley and in the neighboring Serengeti Plateau ...
and the
Aboriginal Australians Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Isl ...
suggest that the sexual division of labor in the Paleolithic was relatively flexible. Men may have participated in gathering plants, firewood and insects, and women may have procured small game animals for consumption and assisted men in driving herds of large game animals (such as woolly mammoths and deer) off cliffs. Additionally, recent research by anthropologist and archaeologist Steven Kuhn from the University of Arizona is argued to support that this division of labor did not exist prior to the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
and was invented relatively recently in human pre-history. Sexual division of labor may have been developed to allow humans to acquire food and other resources more efficiently. Possibly there was approximate parity between men and women during the Middle and Upper Paleolithic, and that period may have been the most gender-equal time in human history.Museum of Antiquites web site
. Retrieved February 13, 2008.
Archaeological evidence from art and funerary rituals indicates that a number of individual women enjoyed seemingly high status in their communities, and it is likely that both sexes participated in decision making. The earliest known Paleolithic
shaman Shamanism is a religious practice that involves a practitioner (shaman) interacting with what they believe to be a spirit world through altered states of consciousness, such as trance. The goal of this is usually to direct spirits or spir ...
( BP) was female.Tedlock, Barbara. 2005. The Woman in the Shaman's Body: Reclaiming the Feminine in Religion and Medicine. New York: Bantam.
Jared Diamond Jared Mason Diamond (born September 10, 1937) is an American geographer, historian, ornithologist, and author best known for his popular science books '' The Third Chimpanzee'' (1991); ''Guns, Germs, and Steel'' (1997, awarded a Pulitzer Priz ...
suggests that the status of women declined with the adoption of agriculture because women in farming societies typically have more pregnancies and are expected to do more demanding work than women in hunter-gatherer societies. Like most modern hunter-gatherer societies, Paleolithic and Mesolithic groups probably followed a largely ambilineal approach. At the same time, depending on the society, the residence could be virilocal, uxorilocal, and sometimes the spouses could live with neither the husband's relatives nor the wife's relatives at all. Taken together, most likely, the lifestyle of hunter-gatherers can be characterized as multilocal.


Sculpture and painting

Early examples of artistic expression, such as the
Venus of Tan-Tan The Venus of Tan-Tan (supposedly, 500,000-300,000 BP) is an alleged artifact found in Morocco. It and its contemporary, the Venus of Berekhat Ram, have been claimed as the earliest representations of the human form. Description The Venus of T ...
and the patterns found on
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae ...
bones from Bilzingsleben in
Thuringia Thuringia (; german: Thüringen ), officially the Free State of Thuringia ( ), is a state of central Germany, covering , the sixth smallest of the sixteen German states. It has a population of about 2.1 million. Erfurt is the capital and lar ...
, may have been produced by Acheulean tool users such as '' Homo erectus'' prior to the start of the Middle Paleolithic period. However, the earliest undisputed evidence of art during the Paleolithic comes from Middle Paleolithic/ Middle Stone Age sites such as Blombos Cave–South Africa–in the form of
bracelet A bracelet is an article of jewellery that is worn around the wrist. Bracelets may serve different uses, such as being worn as an ornament. When worn as ornaments, bracelets may have a supportive function to hold other items of decoration, suc ...
s,
bead A bead is a small, decorative object that is formed in a variety of shapes and sizes of a material such as stone, bone, shell, glass, plastic, wood, or pearl and with a small hole for threading or stringing. Beads range in size from under ...
s, rock art, and ochre used as body paint and perhaps in ritual. Undisputed evidence of art only becomes common in the Upper Paleolithic. Lower Paleolithic
Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...
tool users, according to Robert G. Bednarik, began to engage in symbolic behavior such as art around 850,000 BP. They decorated themselves with beads and collected exotic stones for aesthetic, rather than utilitarian qualities. According to him, traces of the pigment ochre from late Lower Paleolithic Acheulean archaeological sites suggests that Acheulean societies, like later Upper Paleolithic societies, collected and used ochre to create rock art. Nevertheless, it is also possible that the ochre traces found at Lower Paleolithic sites is naturally occurring.
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
humans produced works of art such as cave paintings, Venus figurines, animal carvings, and rock paintings. Upper Paleolithic art can be divided into two broad categories: figurative art such as cave paintings that clearly depicts animals (or more rarely humans); and nonfigurative, which consists of shapes and symbols. Cave paintings have been interpreted in a number of ways by modern archaeologists. The earliest explanation, by the prehistorian Abbe Breuil, interpreted the paintings as a form of magic designed to ensure a successful hunt. However, this hypothesis fails to explain the existence of animals such as
saber-toothed cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million ...
s and lions, which were not hunted for food, and the existence of half-human, half-animal beings in cave paintings. The anthropologist
David Lewis-Williams James David Lewis-Williams (born 1934) is a South African archaeologist. He is best known for his research on southern African San (Bushmen) rock art, of which it can be said that he found a 'Rosetta Stone'. He is the founder and previous direct ...
has suggested that Paleolithic cave paintings were indications of
shaman Shamanism is a religious practice that involves a practitioner (shaman) interacting with what they believe to be a spirit world through altered states of consciousness, such as trance. The goal of this is usually to direct spirits or spir ...
istic practices, because the paintings of half-human, half-animal figures and the remoteness of the caves are reminiscent of modern hunter-gatherer shamanistic practices. Symbol-like images are more common in Paleolithic cave paintings than are depictions of animals or humans, and unique symbolic patterns might have been trademarks that represent different
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
ethnic groups.
Venus figurines A Venus figurine is any Upper Palaeolithic statuette portraying a woman, usually carved in the round.Fagan, Brian M., Beck, Charlotte, "Venus Figurines", ''The Oxford Companion to Archaeology'', 1996, Oxford University Press, pp. 740–741 Most ...
have evoked similar controversy. Archaeologists and anthropologists have described the figurines as representations of goddesses,
pornographic Pornography (often shortened to porn or porno) is the portrayal of sexual subject matter for the exclusive purpose of sexual arousal. Primarily intended for adults,
imagery, apotropaic amulets used for sympathetic magic, and even as self-portraits of women themselves. R. Dale GuthrieR. Dale Guthrie, ''The Nature of Paleolithic Art''. University of Chicago Press, 2006.
Preface
has studied not only the most artistic and publicized paintings, but also a variety of lower-quality art and figurines, and he identifies a wide range of skill and ages among the artists. He also points out that the main themes in the paintings and other artifacts (powerful beasts, risky hunting scenes and the over-sexual representation of women) are to be expected in the fantasies of adolescent males during the Upper Paleolithic. The "Venus" figurines have been theorized, not universally, as representing a mother goddess; the abundance of such female imagery has inspired the theory that religion and society in Paleolithic (and later Neolithic) cultures were primarily interested in, and may have been directed by, women. Adherents of the theory include archaeologist
Marija Gimbutas Marija Gimbutas ( lt, Marija Gimbutienė, ; January 23, 1921 – February 2, 1994) was a Lithuanian archaeologist and anthropologist known for her research into the Neolithic and Bronze Age cultures of " Old Europe" and for her Kurgan hypothesis ...
and feminist scholar
Merlin Stone Merlin Stone (born Marilyn Jacobson, September 27, 1931 – February 23, 2011) was an American author, artist and academic. She was an important thinker of the feminist theology and Goddess movements and is known for her book ''When God Was a ...
, the author of the 1976 book '' When God Was a Woman''. Other explanations for the purpose of the figurines have been proposed, such as Catherine McCoid and LeRoy McDermott's hypothesis that they were self-portraits of woman artists and R.Dale Gutrie's hypothesis that served as "stone age pornography".


Music

The origins of music during the Paleolithic are unknown. The earliest forms of music probably did not use musical instruments other than the human voice or natural objects such as rocks. This early music would not have left an archaeological footprint. Music may have developed from rhythmic sounds produced by daily chores, for example, cracking open nuts with stones. Maintaining a rhythm while working may have helped people to become more efficient at daily activities. An alternative theory originally proposed by
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
explains that music may have begun as a hominin mating strategy. Bird and other animal species produce music such as calls to attract mates. This hypothesis is generally less accepted than the previous hypothesis, but nonetheless provides a possible alternative.
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
(and possibly Middle Paleolithic)Nelson, D.E., ''Radiocarbon dating of bone and charcoal from Divje babe I cave'', cited by Morley, p. 47 humans used flute-like bone pipes as musical instruments,Bahn, Paul (1996) "The atlas of world archeology" Copyright 2000 The Brown Reference Group PLC and music may have played a large role in the religious lives of Upper Paleolithic hunter-gatherers. As with modern hunter-gatherer societies, music may have been used in ritual or to help induce
trance Trance is a state of semi-consciousness in which a person is not self-aware and is either altogether unresponsive to external stimuli (but nevertheless capable of pursuing and realizing an aim) or is selectively responsive in following the dir ...
s. In particular, it appears that animal skin drums may have been used in religious events by Upper Paleolithic shamans, as shown by the remains of drum-like instruments from some Upper Paleolithic graves of shamans and the
ethnographic Ethnography (from Greek ''ethnos'' "folk, people, nation" and ''grapho'' "I write") is a branch of anthropology and the systematic study of individual cultures. Ethnography explores cultural phenomena from the point of view of the subject ...
record of contemporary hunter-gatherer shamanic and ritual practices.


Religion and beliefs

According to James B. Harrod humankind first developed
religious Religion is usually defined as a social- cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatur ...
and spiritual beliefs during the Middle Paleolithic or
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
. Controversial scholars of prehistoric religion and anthropology, James Harrod and Vincent W. Fallio, have recently proposed that religion and spirituality (and art) may have first arisen in Pre-Paleolithic chimpanzees or Early Lower Paleolithic (
Oldowan The Oldowan (or Mode I) was a widespread stone tool archaeological industry (style) in prehistory. These early tools were simple, usually made with one or a few flakes chipped off with another stone. Oldowan tools were used during the Lower ...
) societies. According to Fallio, the common ancestor of chimpanzees and humans experienced altered states of consciousness and partook in ritual, and ritual was used in their societies to strengthen social bonding and group cohesion. Middle Paleolithic humans' use of burials at sites such as Krapina, Croatia ( BP) and
Qafzeh Mount Precipice ( he, הר הקפיצה, "''Har HaKfitsa''"; ar, جبل القفزة, "''Jebel al-Qafzeh''", "Mount of the Leap"), also known as Mount of Precipitation, Mount of the Leap of the Lord and Mount Kedumim is located just outside the ...
, Israel ( BP) have led some anthropologists and archaeologists, such as
Philip Lieberman Philip Lieberman (October 25, 1934 – July 12, 2022) was a cognitive scientist at Brown University, Providence, Rhode Island, United States. Originally trained in phonetics, he wrote a dissertation on intonation. His career focused on topic ...
, to believe that Middle Paleolithic humans may have possessed a belief in an afterlife and a "concern for the dead that transcends daily life". Cut marks on Neanderthal bones from various sites, such as Combe-Grenal and Abri Moula in France, suggest that the Neanderthals—like some contemporary human cultures—may have practiced ritual defleshing for (presumably) religious reasons. According to recent archaeological findings from ''Homo heidelbergensis'' sites in Atapuerca, humans may have begun burying their dead much earlier, during the late Lower Paleolithic; but this theory is widely questioned in the scientific community. Likewise, some scientists have proposed that Middle Paleolithic societies such as Neanderthal societies may also have practiced the earliest form of
totemism A totem (from oj, ᑑᑌᒼ, italics=no or '' doodem'') is a spirit being, sacred object, or symbol that serves as an emblem of a group of people, such as a family, clan, lineage, or tribe, such as in the Anishinaabe clan system. While ''the w ...
or animal worship, in addition to their (presumably religious) burial of the dead. In particular, Emil Bächler suggested (based on archaeological evidence from Middle Paleolithic caves) that a bear cult was widespread among Middle Paleolithic
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While the ...
s. A claim that evidence was found for Middle Paleolithic animal worship  BCE originates from the Tsodilo Hills in the African Kalahari desert has been denied by the original investigators of the site. Animal cults in the Upper Paleolithic, such as the bear cult, may have had their origins in these hypothetical Middle Paleolithic animal cults. Animal worship during the Upper Paleolithic was intertwined with hunting rites. For instance, archaeological evidence from art and bear remains reveals that the bear cult apparently involved a type of sacrificial bear ceremonialism, in which a bear was shot with arrows, finished off by a shot or thrust in the lungs, and ritually worshipped near a clay bear statue covered by a bear fur with the skull and the body of the bear buried separately. Barbara Ehrenreich controversially theorizes that the sacrificial hunting rites of the Upper Paleolithic (and by extension Paleolithic cooperative big-game hunting) gave rise to war or warlike raiding during the following Epipaleolithic and Mesolithic or late Upper Paleolithic. The existence of anthropomorphic images and half-human, half-animal images in the Upper Paleolithic may further indicate that
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
humans were the first people to believe in a pantheon of gods or supernatural beings, though such images may instead indicate shamanistic practices similar to those of contemporary tribal societies. The earliest known undisputed burial of a shaman (and by extension the earliest undisputed evidence of shamans and shamanic practices) dates back to the early
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
era ( BP) in what is now the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
. However, during the early Upper Paleolithic it was probably more common for all members of the band to participate equally and fully in religious ceremonies, in contrast to the religious traditions of later periods when religious authorities and part-time ritual specialists such as shamans, priests and medicine men were relatively common and integral to religious life. Religion was possibly
apotropaic Apotropaic magic (from Greek "to ward off") or protective magic is a type of magic intended to turn away harm or evil influences, as in deflecting misfortune or averting the evil eye. Apotropaic observances may also be practiced out of superst ...
; specifically, it may have involved sympathetic magic. The
Venus figurines A Venus figurine is any Upper Palaeolithic statuette portraying a woman, usually carved in the round.Fagan, Brian M., Beck, Charlotte, "Venus Figurines", ''The Oxford Companion to Archaeology'', 1996, Oxford University Press, pp. 740–741 Most ...
, which are abundant in the Upper Paleolithic archaeological record, provide an example of possible Paleolithic sympathetic magic, as they may have been used for ensuring success in hunting and to bring about fertility of the land and women. The Upper Paleolithic Venus figurines have sometimes been explained as depictions of an
earth goddess An Earth goddess is a deification of the Earth. Earth goddesses are often associated with the "chthonic" deities of the underworld. Ki and Ninhursag are Mesopotamian earth goddesses. In Greek mythology, the Earth is personified as Gaia, corres ...
similar to Gaia, or as representations of a goddess who is the ruler or mother of the animals. James Harrod has described them as representative of female (and male) shamanistic spiritual transformation processes.


Diet and nutrition

Paleolithic hunting and gathering people ate varying proportions of vegetables (including tubers and roots), fruit, seeds (including nuts and wild grass seeds) and insects, meat, fish, and shellfish. However, there is little direct evidence of the relative proportions of plant and animal foods. Although the term "
paleolithic diet The Paleolithic diet, Paleo diet, caveman diet, or stone-age diet is a modern fad diet consisting of foods thought by its proponents to mirror those eaten by humans during the Paleolithic era. The diet avoids processed food and typically incl ...
", without references to a specific timeframe or locale, is sometimes used with an implication that most humans shared a certain diet during the entire era, that is not entirely accurate. The Paleolithic was an extended period of time, during which multiple technological advances were made, many of which had impact on human dietary structure. For example, humans probably did not possess the control of fire until the Middle Paleolithic, or tools necessary to engage in extensive
fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques inclu ...
. On the other hand, both these technologies are generally agreed to have been widely available to humans by the end of the Paleolithic (consequently, allowing humans in some regions of the planet to rely heavily on fishing and hunting). In addition, the Paleolithic involved a substantial geographical expansion of human populations. During the Lower Paleolithic, ancestors of modern humans are thought to have been constrained to Africa east of the
Great Rift Valley The Great Rift Valley is a series of contiguous geographic trenches, approximately in total length, that runs from Lebanon in Asia to Mozambique in Southeast Africa. While the name continues in some usages, it is rarely used in geology as it ...
. During the Middle and Upper Paleolithic, humans greatly expanded their area of settlement, reaching ecosystems as diverse as
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
and
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S. ...
, and adapting their diets to whatever local resources were available. Another view is that until the Upper Paleolithic, humans were frugivores (fruit eaters) who supplemented their meals with carrion, eggs, and small prey such as baby birds and mussels, and only on rare occasions managed to kill and consume big game such as antelopes. This view is supported by studies of higher apes, particularly
chimpanzees The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
. Chimpanzees are the closest to humans genetically, sharing more than 96% of their DNA code with humans, and their digestive tract is functionally very similar to that of humans. Chimpanzees are primarily frugivores, but they could and would consume and digest animal flesh, given the opportunity. In general, their actual diet in the wild is about 95% plant-based, with the remaining 5% filled with insects, eggs, and baby animals. In some ecosystems, however, chimpanzees are predatory, forming parties to hunt monkeys. Some comparative studies of human and higher primate digestive tracts do suggest that humans have evolved to obtain greater amounts of calories from sources such as animal foods, allowing them to shrink the size of the gastrointestinal tract relative to body mass and to increase the brain mass instead. Anthropologists have diverse opinions about the proportions of plant and animal foods consumed. Just as with still existing hunters and gatherers, there were many varied "diets" in different groups, and also varying through this vast amount of time. Some paleolithic hunter-gatherers consumed a significant amount of meat and possibly obtained most of their food from hunting, while others were believed to have a primarily plant-based diet. Most, if not all, are believed to have been opportunistic omnivores. One hypothesis is that carbohydrate
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growin ...
s (plant underground storage organs) may have been eaten in high amounts by pre-agricultural humans. It is thought that the Paleolithic diet included as much as per day of fruit and vegetables. The relative proportions of plant and animal foods in the diets of Paleolithic people often varied between regions, with more meat being necessary in colder regions (which were not populated by anatomically modern humans until  BP).) It is generally agreed that many modern hunting and fishing tools, such as fish hooks, nets, bows, and poisons, were not introduced until the Upper Paleolithic and possibly even Neolithic. The only hunting tools widely available to humans during any significant part of the Paleolithic were hand-held spears and harpoons. There is evidence of Paleolithic people killing and eating seals and elands as far as  BP. On the other hand, buffalo bones found in African caves from the same period are typically of very young or very old individuals, and there is no evidence that pigs, elephants, or rhinos were hunted by humans at the time. Paleolithic peoples suffered less
famine A famine is a widespread scarcity of food, caused by several factors including war, natural disasters, crop failure, population imbalance, widespread poverty, an economic catastrophe or government policies. This phenomenon is usually accompani ...
and
malnutrition Malnutrition occurs when an organism gets too few or too many nutrients, resulting in health problems. Specifically, it is "a deficiency, excess, or imbalance of energy, protein and other nutrients" which adversely affects the body's tissues ...
than the Neolithic farming tribes that followed them.p. 2
/ref> This was partly because Paleolithic hunter-gatherers accessed a wider variety of natural foods, which allowed them a more nutritious diet and a decreased risk of famine. Many of the famines experienced by Neolithic (and some modern) farmers were caused or amplified by their dependence on a small number of crops. It is thought that wild foods can have a significantly different nutritional profile than cultivated foods. The greater amount of meat obtained by hunting big game animals in Paleolithic diets than Neolithic diets may have also allowed Paleolithic hunter-gatherers to enjoy a more nutritious diet than Neolithic agriculturalists. It has been argued that the shift from hunting and gathering to agriculture resulted in an increasing focus on a limited variety of foods, with meat likely taking a back seat to plants. It is also unlikely that Paleolithic hunter-gatherers were affected by modern diseases of affluence such as
type 2 diabetes Type 2 diabetes, formerly known as adult-onset diabetes, is a form of diabetes mellitus that is characterized by high blood sugar, insulin resistance, and relative lack of insulin. Common symptoms include increased thirst, frequent urinatio ...
,
coronary heart disease Coronary artery disease (CAD), also called coronary heart disease (CHD), ischemic heart disease (IHD), myocardial ischemia, or simply heart disease, involves the reduction of blood flow to the heart muscle due to build-up of atherosclerotic pl ...
, and
cerebrovascular disease Cerebrovascular disease includes a variety of medical conditions that affect the blood vessels of the brain and the cerebral circulation. Arteries supplying oxygen and nutrients to the brain are often damaged or deformed in these disorders. Th ...
, because they ate mostly lean meats and plants and frequently engaged in intense physical activity, and because the average lifespan was shorter than the age of common onset of these conditions. Large-seeded legumes were part of the human diet long before the Neolithic Revolution, as evident from archaeobotanical finds from the
Mousterian The Mousterian (or Mode III) is an archaeological industry of stone tools, associated primarily with the Neanderthals in Europe, and to the earliest anatomically modern humans in North Africa and West Asia. The Mousterian largely defines the l ...
layers of Kebara Cave, in Israel. There is evidence suggesting that Paleolithic societies were gathering wild cereals for food use at least as early as 30,000 years ago. However, seeds—such as grains and beans—were rarely eaten and never in large quantities on a daily basis. Recent archaeological evidence also indicates that
winemaking Winemaking or vinification is the production of wine, starting with the selection of the fruit, its fermentation into alcohol, and the bottling of the finished liquid. The history of wine-making stretches over millennia. The science of wine and ...
may have originated in the Paleolithic, when early humans drank the juice of naturally fermented wild grapes from animal-skin pouches. Paleolithic humans consumed animal organ meats, including the
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
s,
kidney The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; blo ...
s, and
brain A brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It is located in the head, usually close to the sensory organs for senses such as vision. It is the most complex organ in a ve ...
s. Upper Paleolithic cultures appear to have had significant knowledge about plants and herbs and may have sometimes practiced rudimentary forms of
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
. In particular, bananas and
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growin ...
s may have been cultivated as early as 25,000 BP in
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainlan ...
. In the Paleolithic Levant, 23,000 years ago, cereals cultivation of
emmer Emmer wheat or hulled wheat is a type of awned wheat. Emmer is a tetraploid (4''n'' = 4''x'' = 28 chromosomes). The domesticated types are ''Triticum turgidum'' subsp. ''dicoccum'' and ''Triticum turgidum ''conv.'' durum''. The wild plant is ...
,
barley Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley p ...
, and
oats The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural, unlike other cereals and pseudocereals). While oats are suitable for human co ...
has been observed near the Sea of Galilee. Late Upper Paleolithic societies also appear to have occasionally practiced
pastoralism Pastoralism is a form of animal husbandry where domesticated animals (known as " livestock") are released onto large vegetated outdoor lands (pastures) for grazing, historically by nomadic people who moved around with their herds. The a ...
and animal husbandry, presumably for dietary reasons. For instance, some European late Upper Paleolithic cultures domesticated and raised
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 sub ...
, presumably for their meat or milk, as early as 14,000 BP. Humans also probably consumed
hallucinogenic Hallucinogens are a large, diverse class of psychoactive drugs that can produce altered states of consciousness characterized by major alterations in thought, mood, and perception as well as other changes. Most hallucinogens can be categorized ...
plants during the Paleolithic. The
Aboriginal Australians Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Isl ...
have been consuming a variety of native animal and plant foods, called
bushfood Bush tucker, also called bush food, is any food native to Australia and used as sustenance by Indigenous Australians, the Aboriginal and Torres Strait Islander peoples, but it can also describe any native flora or fauna used for culinary or ...
, for an estimated 60,000 years, since the Middle Paleolithic. In February 2019, scientists reported evidence, based on
isotope Isotopes are two or more types of atoms that have the same atomic number (number of protons in their nuclei) and position in the periodic table (and hence belong to the same chemical element), and that differ in nucleon numbers (mass numb ...
studies, that at least some Neanderthals may have eaten meat. People during the Middle Paleolithic, such as the Neanderthals and Middle Paleolithic Homo sapiens in Africa, began to catch shellfish for food as revealed by shellfish cooking in Neanderthal sites in Italy about 110,000 years ago and in Middle Paleolithic ''Homo sapiens'' sites at
Pinnacle Point Pinnacle Point a small promontory immediately south of Mossel Bay, a town on the southern coast of South Africa. Excavations since the year 2000 of a series of caves at Pinnacle Point have revealed occupation by Middle Stone Age people between ...
, South Africa around 164,000 BP. Although fishing only became common during the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
,
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
have been part of human diets long before the dawn of the Upper Paleolithic and have certainly been consumed by humans since at least the Middle Paleolithic. For example, the Middle Paleolithic ''Homo sapiens'' in the region now occupied by the
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
hunted large -long
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive ...
with specialized barbed fishing points as early as 90,000 years ago. The invention of fishing allowed some Upper Paleolithic and later hunter-gatherer societies to become sedentary or semi-nomadic, which altered their social structures. Example societies are the
Lepenski Vir Lepenski Vir ( sr-cyr, Лепенски Вир, "Lepena Whirlpool"), located in Serbia, is an important archaeological site of the Mesolithic Iron Gates culture of the Balkans. The latest radiocarbon and AMS data suggests that the chronology of ...
as well as some contemporary hunter-gatherers, such as the
Tlingit The Tlingit ( or ; also spelled Tlinkit) are indigenous peoples of the Pacific Northwest Coast of North America. Their language is the Tlingit language (natively , pronounced ),
. In some instances (at least the Tlingit), they developed social stratification,
slavery Slavery and enslavement are both the state and the condition of being a slave—someone forbidden to quit one's service for an enslaver, and who is treated by the enslaver as property. Slavery typically involves slaves being made to perf ...
, and complex social structures such as
chiefdom A chiefdom is a form of hierarchical political organization in non-industrial societies usually based on kinship, and in which formal leadership is monopolized by the legitimate senior members of select families or 'houses'. These elites form a ...
s. Anthropologists such as Tim White suggest that cannibalism was common in human societies prior to the beginning of the Upper Paleolithic, based on the large amount of “butchered human" bones found in Neanderthal and other Lower/Middle Paleolithic sites. Cannibalism in the Lower and Middle Paleolithic may have occurred because of food shortages. However, it may have been for religious reasons, and would coincide with the development of religious practices thought to have occurred during the Upper Paleolithic. Nonetheless, it remains possible that Paleolithic societies never practiced cannibalism, and that the damage to recovered human bones was either the result of
excarnation In archaeology and anthropology, the term excarnation (also known as defleshing) refers to the practice of removing the flesh and organs of the dead before burial. Excarnation may be achieved through natural means, such as leaving a dead body expo ...
or
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
by carnivores such as
saber-toothed cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million ...
s, lions, and hyenas. A modern-day diet known as the
Paleolithic diet The Paleolithic diet, Paleo diet, caveman diet, or stone-age diet is a modern fad diet consisting of foods thought by its proponents to mirror those eaten by humans during the Paleolithic era. The diet avoids processed food and typically incl ...
exists, based on restricting consumption to the foods presumed to be available to anatomically modern humans prior to the advent of settled
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
.


See also

* Abbassia Pluvial *
Bontnewydd Palaeolithic site The Bontnewydd palaeolithic site (), also known in its unmutated form as Pontnewydd (Welsh language: 'New bridge'), is an archaeological site near St Asaph, Denbighshire, Wales. It is one of only three sites in Britain to have produced fossils ...
*
Caveman The caveman is a stock character representative of primitive humans in the Paleolithic. The popularization of the type dates to the early 20th century, when Neanderthals were influentially described as "simian" or " ape-like" by Marcellin Bo ...
* Japanese Paleolithic *
Lascaux Lascaux ( , ; french: Grotte de Lascaux , "Lascaux Cave") is a network of caves near the village of Montignac, in the department of Dordogne in southwestern France. Over 600 parietal wall paintings cover the interior walls and ceilings of ...
* Last Glacial Maximum * List of Paleolithic archaeological sites by continent *
Luzia Woman Luzia Woman () is the name for an Upper Paleolithic period skeleton of a Paleo-Indian woman who was found in a cave in Brazil. Some archaeologists originally thought the young woman may have been part of a migratory wave of immigrants prior to t ...
*
Mousterian Pluvial The Mousterian Pluvial is a mostly obsolete term for a prehistoric wet and rainy ( pluvial) period in North Africa. It was described as beginning around 50,000 years before the present ( BP), lasting roughly 20,000 years, and ending ca. 30,000 BP. ...
*
Origins of society The origins of society — the evolutionary emergence of distinctively human social organization — is an important topic within evolutionary biology, anthropology, prehistory and palaeolithic archaeology. While little is known for certain, debates ...
*
Palaeoarchaeology Palaeoarchaeology (or paleoarcheology) is the archaeology of deep time. Paleoarchaeologists' studies focus on hominid fossils ranging from 15,000,000 to 10,000 years ago, and human evolution and the ways in which humans have adapted to the environ ...
*
Settlement of the Americas The settlement of the Americas began when Paleolithic hunter-gatherers entered North America from the North Asian Mammoth steppe via the Beringia land bridge, which had formed between northeastern Siberia and western Alaska due to the lowering o ...
*
Turkana Boy Turkana Boy, also called Nariokotome Boy, is the name given to fossil KNM-WT 15000, a nearly complete skeleton of a ''Homo ergaster'' youth who lived 1.5 to 1.6 million years ago. This specimen is the most complete early hominin skeleton ever ...


References


External links


Human Timeline (Interactive)
Smithsonian,
National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...
(August 2016).
Donsmaps: a vast repository of Paleolithic resources

Interactive Timeline Simile/Timemap index of Eurasian sites
{{Authority control Pleistocene Historical eras