The Pacific kingfisher (''Todiramphus sacer'') is a medium-sized

kingfisher Kingfishers are a family, the Alcedinidae, of small to medium-sized, brightly colored birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species found in the tropical regions of Africa, Asia, and Oceania, ...

belonging to the subfamily Halcyoninae, the

tree kingfisher The tree kingfishers, also called wood kingfishers or Halcyoninae, are the most numerous of the three subfamilies of birds in the kingfisher family, with around 70 species divided into 12 genera, including several species of kookaburras. Th ...

s. It has a wide range throughout the South Pacific islands. It was previously considered a subspecies of the collared kingfisher.

Taxonomy

The Pacific kingfisher was formally described in 1788 by the German naturalist

Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich Gmelin Ferdinand Christoph Oetinger , academic_advisors = , docto ...

in his revised and expanded edition of

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...

's ''

Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...

''. He placed it with the other kingfishers in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

Alcedo ''Alcedo'' is a genus of birds in the kingfisher subfamily Alcedininae. The genus was introduced by Carl Linnaeus in 1758 in the 10th edition of his ''Systema Naturae''. The type species is the common kingfisher (''Alcedo ispida'', now ''Alcedo ...

'' and coined the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...

''Alcedo sacra''. Gmelin based his description on the "Sacred kingfisher" that had been described and illustrated in 1782 by the English ornithologist John Latham in his multi-volume ''A General Synopsis of Birds''. Latham had access to a specimen in the

Leverian Museum The Leverian collection was a natural history and ethnographic collection assembled by Ashton Lever. It was noted for the content it acquired from the voyages of Captain James Cook. For three decades it was displayed in London, being broken up ...

in London that had been collected during

James Cook James Cook (7 November 1728 Old Style date: 27 October – 14 February 1779) was a British explorer, navigator, cartographer, and captain in the British Royal Navy, famous for his three voyages between 1768 and 1779 in the Pacific Ocean and ...

's third voyage to the Pacific Ocean from 1776 to 1780. The type locality was designated as the island of Tongatabu by

Alexander Wetmore Frank Alexander Wetmore (June 18, 1886 – December 7, 1978) was an American ornithologist and avian paleontologist. He was the sixth Secretary of the Smithsonian Institution. Early life and education The son of a Country Physician, Frank Ale ...

in 1919. The Pacific kingfisher is now one of 30 species placed in the genus '' Todiramphus'' that was introduced in 1827 by

René Lesson René-Primevère Lesson (20 March 1794 – 28 April 1849) was a French surgeon, naturalist, ornithologist, and herpetologist. Biography Lesson was born at Rochefort, and entered the Naval Medical School in Rochefort at the age of sixteen. H ...

. The word ''Todiramphus'' combines the genus name '' Todus'' with the

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...

''rhamphos'' meaning "bill". The specific epithet ''sacer'' is

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

meaning "sacred". The Pacific kingfisher was formerly considered as a

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...

of the collared kingfisher (''Todiramphus chloris''). A

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study published in 2015 found that ''T. chloris'' was

paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ...

. In the changes to create monophyletic species, ''T. sacer'' was promoted to species status and many of the ''T. chloris'' subspecies were moved to ''T. sacer''.

Subspecies

There are 22 recognised subspecies: ;Vanuatu * ''T. s. torresianus'' ( Mayr, 1931) – Hiw and Lo * ''T. s. santoensis'' (Mayr, 1931) –

Banks Islands The Banks Islands (in Bislama ''Bankis'') are a group of islands in northern Vanuatu. Together with the Torres Islands to their northwest, they make up the northernmost province of Torba. The island group lies about north of Maewo, and inclu ...

southwards to Espiritu Santo and Malo * ''T. s. juliae'' (

Heine Heine is both a surname and a given name of German origin. People with that name include: People with the surname * Albert Heine (1867–1949), German actor * Alice Heine (1858–1925), American-born princess of Monaco * Armand Heine (1818–188 ...

, 1860) – Aoba and Maewo southwards to

Efate Efate (french: Éfaté) is an island in the Pacific Ocean which is part of the Shefa Province in Vanuatu. It is also known as Île Vate. Geography It is the most populous (approx. 66,000) island in Vanuatu. Efate's land area of makes it Vanu ...

* ''T. s. erromangae'' (Mayr, 1938) –

Erromango Erromango is the fourth largest island in the Vanuatu archipelago. With a land area of it is the largest island in Tafea Province, the southernmost of Vanuatu's six administrative regions. Name The endonym for Erromango in Erromangan is ''Nel ...

and Anatom * ''T. s. tannensis'' ( Sharpe, 1892) – Tanna ;Solomon Islands * ''T. s. pavuvu'' (Mayr, 1935) – Pavuvu * ''T. s. mala'' (Mayr, 1935) –

Malaita Malaita is the primary island of Malaita Province in Solomon Islands. Malaita is the most populous island of the Solomon Islands, with a population of 161,832 as of 2021, or more than a third of the entire national population. It is also the se ...

* ''T. s. solomonis'' ( Ramsay, EP, 1882) –

Makira The island of Makira (also known as San Cristobal and San Cristóbal) is the largest island of Makira-Ulawa Province in the Solomon Islands. It is third most populous island after Malaita and Guadalcanal, with a population of 55,126 as of 2020. ...

and adjacent islands * ''T. s. sororum'' (Galbraith, ICJ & Galbraith, EH, 1962) – Malaupaina and Malaulalo * ''T. s. amoenus'' (Mayr, 1931) – Rennell and

Bellona Bellona may refer to: Places *Bellona, Campania, a ''comune'' in the Province of Caserta, Italy * Bellona Reef, a reef in New Caledonia *Bellona Island, an island in Rennell and Bellona Province, Solomon Islands Ships * HMS ''Bellona'' (1760), a 7 ...

* ''T. s. ornatus'' (Mayr, 1931) – Nendo and Tinakula * ''T. s. brachyurus'' (Mayr, 1931) –

Reef Islands The Reef Islands are a loose collection of 16 islands in the northwestern part of the Solomon Islands province of Temotu. These islands have historically also been known by the names of Swallow Islands and Matema Islands. Geography The islan ...

* ''T. s. vicina'' (Mayr, 1931) –

Duff Islands The Duff Islands are a small island group lying to the northeast of the Santa Cruz Islands in the Solomon Islands province of Temotu. They are also sometimes known as the Wilson Islands. Location and geography The islands are located at 9°5 ...

* ''T. s. utupuae'' (Mayr, 1931) –

Utupua Utupua is an island in the Santa Cruz Islands, located 66 km to the Southeast of the main Santa Cruz group, between Vanikoro and Santa Cruz proper (Nendo Island). This island belongs administratively to the Temotu Province of the Solomon Is ...

* ''T. s. melanodera'' (Mayr, 1931) –

Vanikoro Vanikoro (sometimes wrongly named ''Vanikolo'') is an island in the Santa Cruz group, located to the Southeast of the main Santa Cruz group. It is part of the Temotu Province of the Solomon Islands. The name ''Vanikoro'' is always used as thou ...

;Fiji * ''T. s. vitiensis'' ( Peale, 1849) –

Vanua Levu Vanua Levu (pronounced ), formerly known as Sandalwood Island, is the second largest island of Fiji. Located to the north of the larger Viti Levu, the island has an area of and a population of 135,961 . Geology Fiji lies in a tectonically ...

Taveuni Taveuni (pronounced ) is the third-largest island in Fiji, after Viti Levu and Vanua Levu, with a total land area of . The cigar-shaped island, a massive shield volcano which rises from the floor of the Pacific Ocean, is situated to the east of ...

Viti Levu Viti Levu (pronounced ) is the largest island in the Republic of Fiji. It is the site of the nation's capital, Suva, and home to a large majority of Fiji's population. Geology Fiji lies in a tectonically complex area between the Australian ...

, Koro, Ovalau and Gau * ''T. s. marinus'' (Mayr, 1941) – Lau Archipelago * ''T. s. eximius'' (Mayr, 1941) –

Kadavu Kadavu may refer to: * Kadavu Island, the fourth largest island in Fiji * Kadavu Group, an archipelago in Fiji including Kadavu Island * Kadavu Province Kadavu Province is one of fourteen provinces of Fiji, and forms part of the Eastern Division ...

;Polynesia * ''T. s. regina'' (Mayr, 1941) – Futuna * ''T. s. pealei'' ( Finsch &

Hartlaub Karel Johan Gustav Hartlaub (8 November 1814 – 29 November 1900) was a German physician and ornithologist. Hartlaub was born in Bremen, and studied at Bonn and Berlin before graduating in medicine at Göttingen. In 1840, he began to study and c ...

, 1867) – Tutuila * ''T. s. manuae'' (Mayr, 1941) –

Ofu-Olosega Ofu and Olosega are parts of a volcanic doublet in the Manuʻa Islands, which is a part of American Samoa in the Samoan Islands. These twin islands, formed from shield volcanoes, have a combined length of 6 km and a combined area of . Togethe ...

and Tau * ''T. s. sacer'' ( Gmelin, JF, 1788) – central and southern

Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...

Description

It has a variety of

calls Call or Calls may refer to: Arts, entertainment, and media Games * Call, a type of betting in poker * Call, in the game of contract bridge, a bid, pass, double, or redouble in the bidding stage Music and dance * Call (band), from Lahore, Pak ...

which vary geographically. The most typical call is loud, harsh and metallic and is repeated several times.

Distribution and habitat

It is most commonly found in coastal areas, particularly in

mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...

swamps. It also inhabits farmland, open woodland, grassland and gardens. It is usually seen further inland than the collared kingfisher, where it was previously considered conspecific, ranging into forest or into mountain areas. Birds often perch conspicuously on wires, rocks or bare branches. On the

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...

islands it is usually common in a variety of coastal and inland habitats with various subspecies present on the

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...

Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of ...

Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consis ...

, and

American Samoa American Samoa ( sm, Amerika Sāmoa, ; also ' or ') is an unincorporated territory of the United States located in the South Pacific Ocean, southeast of the island country of Samoa. Its location is centered on . It is east of the Internatio ...

Behaviour and ecology

Food and feeding

Small

crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...

s are the favoured food in coastal regions but a wide variety of other animals are eaten including

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...

worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...

snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class ...

shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are ref ...

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...

s, small

fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...

and sometimes other smal
birds
as well. The bird perches almost motionless for long periods waiting for prey. When it spots something it glides down to catch it and then flies back to the perch where larger items are pounded against the branch to subdue them. Any indigestible remains are regurgitated as pellets.

Breeding

The

nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...

is a hole, either a natural tree hole or a burrow excavated by the birds themselves in a rotten tree,

termite Termites are small insects that live in colonies and have distinct castes ( eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blat ...

mound or earth bank. They will also occupy old

woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions ...

holes. Two to seven rounded whitish eggs are laid directly on the floor of the burrow with no nest material used. Both parents take part in incubating the eggs and feeding the chicks. The young birds leave the nest about 44 days after hatching. Two broods are often raised in a year.

References

External links

* * {{Taxonbar, from=Q22285209 Pacific kingfisher Birds of the Pacific Ocean Pacific kingfisher Pacific kingfisher