The Pacific goliath grouper (''Epinephelus quinquefasciatus''), also known as the Pacific itajara grouper, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

, a

grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...

from the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...

which is part of the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some case ...

, which also includes the

anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...

and sea basses. It found in the eastern Pacific Ocean where it is associated with reefs. It is related to the Atlantic goliath grouper.

Description

The Pacific goliath grouper has a robust, oblong body. The area between the eyes is wide and flat. The margin of the preopercle has fine serrations and an angled edge. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

contains 11 spines and 15–16 soft rays while the

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

has 3 spines and 8 soft rays. The

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

istail rounded. The head and body are grey or greenish in colour marked with pale blotches and small dark spots which are scattered over the upper head and body, as well as being on the pectoral fins. Subadult fish which are less than in length are overall greenish to tawny brown in colour with diagonal, irregular darker brown bars on the body and caudal fin. The juveniles have heavy spotting on the head, the soft-rayed part of the dorsal fin and the pectoral,

pelvic The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

, and caudal fins, They have 5 diagonal black bars on the body which reach onto the dorsal and anal fins and there is a black bar on base of the caudal fin. This is one of the largest species of grouper, attaining a maximum

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of .

Distribution

The Pacific goliath grouper is found in the Eastern Pacific Ocean off the west coasts of the Americas. It is found from the

Gulf of California The Gulf of California ( es, Golfo de California), also known as the Sea of Cortés (''Mar de Cortés'') or Sea of Cortez, or less commonly as the Vermilion Sea (''Mar Bermejo''), is a marginal sea of the Pacific Ocean that separates the Baja Ca ...

and the Pacific coast of

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

south to Peru. Its range also includes the islands groups of

Revillagigedos The Revillagigedo Islands ( es, Islas Revillagigedo, ) or Revillagigedo Archipelago are a group of four volcanic islands in the Pacific Ocean, known for their unique ecosystem. They lie approximately from Socorro Island south and southwest of C ...

in Mexico,

Cocos Island Cocos Island ( es, Isla del Coco) is an island in the Pacific Ocean administered by Costa Rica, approximately southwest of the Costa Rican mainland. It constitutes the 11th of the 13 districts of Puntarenas Canton of the Province of Puntarenas ...

in Costa Rica and Malpelo Island in Colombia.

Habitat and biology

The Pacific goliath grouper is found on offshore rocky reefs as adults, although it has also been recorded in inshore areas. The juveniles inhabit mangroves, estuaries, lagoons and bays. It has been recorded feeding on sharks, rays, crustaceans, cephalopods, other fishes and even

sea snake Sea snakes, or coral reef snakes, are elapid snakes that inhabit marine environments for most or all of their lives. They belong to two subfamilies, Hydrophiinae and Laticaudinae. Hydrophiinae also includes Australasian terrestrial snakes, wher ...

s and mammals. Little is known about its biology but it is thought to be similar to the Atlantic goliath grouper.

Taxonomy

The Pacific goliath grouper was first formally described as ''Serranus quinquefasciatus'' in 1868 by the

French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

and

artist An artist is a person engaged in an activity related to creating art, practicing the arts, or demonstrating an art. The common usage in both everyday speech and academic discourse refers to a practitioner in the visual arts only. However, th ...

Marie Firmin Bocourt Marie Firmin Bocourt (19 April 1819 – 4 February 1904) was a French zoologist and artist. As a young man, he worked as a preparateur for the zoologist Gabriel Bibron (1805–1848), later serving as a museum artist. In 1861, he was sent to Thail ...

(1819–1904) with the type locality given as the mouth of Río Nagualate on the Pacific coast of Guatemala. Felipe Poey assigned the giant and goliath groupers (''

Epinephelus itajara The Atlantic goliath grouper or itajara (''Epinephelus itajara''), formerly known as the jewfish, is a saltwater fish of the grouper family and one of the largest species of bony fish. The species can be found in the west ranging from northeaste ...

'' and '' E. quinquefasciatus'', which was then regarded as a

synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

of ''E itajara'') to the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Promicrops'' but in 1972 this was designated as a

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

of ''

Epinephelus ''Epinephelus'' is a genus of marine ray-finned fish, groupers from the subfamily Epinephelinae, part of the family Serranidae, which also includes the anthias and sea basses. They are predatory fish, largely associated with reefs and are found i ...

''. However, these species are still each others closest relatives.

Utilisation

The Pacific goliath grouper is a sought after quarry species for recreational and commercial fisheries in the entirety of its range. It has declined over much of its range, and, in Colombia, smaller goliath groupers of lengths less than are regarded as the most valuable. This leads fishermen to target small and sexually immature groupers which threatens the local survival of the species by taking the fish before they get an opportunity to reproduce.

References

{{Taxonbar, from=Q5501215 Epinephelus Fish of the Pacific Ocean Fish of the Gulf of California Western Central American coastal fauna Fish described in 1868