Northern brown snake
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''Pseudonaja nuchalis'', the northern brown snake or ''gwardar'', is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of very fast, highly
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
elapid Elapidae (, commonly known as elapids ; grc, ἔλλοψ ''éllops'' "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus Emydoce ...
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
native to Australia. Its colour and pattern are rather variable, depending largely on its location. The name ''gwardar'' is a word meaning "go the long way around" in an Aboriginal language . This may be regarded as advice for people who come across the species in the wild: that is, while ''P. nuchalis'' is generally cautious, shy, and inclined to retreat rather than attack, it will defend itself if cornered.


Description

The northern brown snake grows up to in total length (including tail). Its back can feature shades of orange-brown with flecks and bands, or appear plain. Its belly is cream to orange with pink blotches. Some individuals have jet black heads (this can cause it to be confused with the black-headed python), while others feature a black 'V' shape on the back of their neck, below their head.


Distribution and habitat

The northern brown snake has a wide distribution and is found across the north of the Australian continent, including the
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an Australian territory in the central and central northern regions of Australia. The Northern Territory shares its borders with Western Aust ...
,
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
and
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
. The northern brown snake is a ground-dwelling snake that prefers drier
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s, but is also found in coastal
eucalypt Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', '' Stockwellia'', ''Allosyn ...
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s,
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (se ...
s, and
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s. Although the northern brown is not an
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
species, it is not uncommon for it to climb small shrubs or
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s. It also hides in crevices and under rocks, and in urban areas can be found under rubbish or tin piles.


Diet

''Pseudonaja nuchalis''
preys Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
upon small mammals and reptiles, including lizards and mice.


Lifespan and reproduction

Little is known about the northern brown snake's lifespan. Mating season is roughly from September to November and the female usually produces around 11-14 eggs, but may produce up to 38.


Venom and symptoms

Although the northern brown snake's venom is not the most toxic in the brown snake genus, its average delivery contains a relatively high quantity of venom; thus the northern brown snake has high potential to deliver a deadly bite. Its venom contains neurotoxins,
nephrotoxin Nephrotoxicity is toxicity in the kidneys. It is a poisonous effect of some substances, both toxic chemicals and medications, on kidney function. There are various forms, and some drugs may affect kidney function in more than one way. Nephrotoxin ...
s, and a
procoagulant Coagulation, also known as clotting, is the process by which blood changes from a liquid to a gel, forming a blood clot. It potentially results in hemostasis, the cessation of blood loss from a damaged vessel, followed by repair. The mechanism o ...
, although humans are not usually affected by the neurotoxins. The bite is usually painless and difficult to see due to their small fangs. Human symptoms of a northern brown snake bite are headache, nausea/vomiting, abdominal pain, severe coagulopathy, and sometimes kidney damage. In dogs and cats, paralysis is also likely to occur.


Behaviour

The northern brown snake is known to be very aggressive when disturbed or threatened, but like most snakes, usually prefers to retreat from danger. It may develop nocturnal habits during the warmer months, but is otherwise active during the day and enjoys sunlight. The snake has also been known to practise cannibalism, although this is not common. Northern brown snakes kill their prey with a combination of venom and
constriction Constriction is a method used by various snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake i ...
.


References


Further reading

* Boulenger GA (1896). ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., Containing the Colubridæ (Opisthoglyphæ and Proteroglyphæ), ...'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I-XXV. (''Diemenia nuchalis'', pp. 326–327). * Günther A (1858). ''Catalogue of the Colubrine Snakes in the Collection of the British Museum.'' London: Trustees of the British Museum. (Taylor and Francis, printers). xvi+ 281 pp. (''Pseudonaja nuchalis'', new species, pp. 227–228). {{Taxonbar, from=Q2713992 nuchalis Reptiles of Western Australia Reptiles described in 1858 Taxa named by Albert Günther Snakes of Australia