The Neuropterida are a

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

holometabolous Holometabolism, also called complete metamorphosis, is a form of insect development which includes four life stages: egg, larva, pupa, and imago (or adult). Holometabolism is a synapomorphic trait of all insects in the clade Holometabola. Immatur ...

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s, sometimes placed at the

superorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...

level. The clade contains the orders

Neuroptera The insect order (biology), order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera is grouped together with the Megaloptera (alderflies, f ...

(lacewings, antlions),

Megaloptera Megaloptera is an order of insects. It contains the alderflies, dobsonflies and fishflies, and there are about 300 known species. The order's name comes from Ancient Greek, from ''mega-'' (μέγα-) "large" + ''pteryx'' (πτέρυξ) "wi ...

(alderflies, dobsonflies), and

Raphidioptera Snakeflies are a group of predatory insects comprising the order (biology), order Raphidioptera with two extant families: Raphidiidae and Inocelliidae, consisting of roughly 260 species. In the past, the group had a much wider distribution than i ...

(snakeflies), and includes over 5,700 described species. Historically, the name

referred to this entire group, but it now refers only to lacewings and their relatives (antlions), which were formerly known as Planipennia. As part of the

Holometabola Holometabola (from Ancient Greek "complete" + "change"), also known as Endopterygota (from "inner" + "wing" + Neo-Latin "-having"), is a supra-order (biology), ordinal clade of insects within the infraclass Neoptera that go through distincti ...

and related to

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

s, they can be considered an unranked

taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

. Arguably, the Holometabola might instead be considered an unranked clade, and divided into numerous superorders to signify the close relationships of certain holometabolan groups.Haaramo, Mikko (2008): Mikko's Phylogeny Archive
Neuropterida
Version of 11 March 2008. Retrieved 5 May 2008. Some authors formerly included the

Mecoptera Mecoptera (from the Greek language, Greek: ''mecos'' = "long", ''ptera'' = "wings") is an Order (biology), order of insects in the superorder Holometabola with about six hundred species in nine Family (biology), families worldwide. Mecopterans a ...

(scorpionflies) in Neuropterida, but they actually belong to the Mecopteroidea (or Antliophora), the holometabolan clade that also contains

true flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...

and

flea Flea, the common name for the order (biology), order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by hematophagy, ingesting the blood of their hosts. Adult f ...

s. Neuropterida are fairly primitive-looking insects with large wings but weak wing muscles, resulting in clumsy flight. Most adults are active at dusk or night, and many of the

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e are aquatic that live in

river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...

s. The larvae, and in many cases the adults as well, are

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

s of small

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

s. Adult neuropteridans range in size from that of a

midge A midge is any small fly, including species in several family (biology), families of non-mosquito nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid ...

to that of a large

dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threat ...

( wingspan); the largest species tend to resemble drab, clumsily flying

damselflies Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies (which constitute the other odonatan suborder, Epiprocta) but are usually smaller and have slimmer bodies. Most species fold the win ...

. In addition to the three living orders, the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

Rafaelidae are an entirely

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

family of Neuropterida. Their position is indeterminate but probably rather basal; thus, the single

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

'' Rafaeliana'' from the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

Santana Formation's Crato Member in

Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...

might be better placed directly in the Neuropterida for the time being, without assigning it to an order, until relatives are found and/or its systematic position is better resolved. The extinct order Glosselytrodea may also be a member or close relative, though its classification is unclear. A phylogenomic analysis published in 2023 confirmed the topology of the neuropterid orders and found the relationships between the families of Neuropterida as shown in the following phylogenetic tree.

References

External links

* Oswald, John D. (2023)
Neuropterida Species of the World. Lacewing Digital Library, Research Publication No. 1.
(an online catalog of the Neuropterida species of the world) * Oswald, John D. (2023)
Bibliography of the Neuropterida. Lacewing Digital Library, Research Publication No. 2.
(an online bibliography of the global scientific literature of the Neuropterida)
Lacewing Digital Library
(a web portal that provides access to a suite of online resources that contain data on the superorder Neuropterida) {{Taxonbar, from=Q730799 Insect superorders

Neuropterida The Neuropterida are a clade of holometabolous insects, sometimes placed at the superorder level. The clade contains the orders Neuroptera (lacewings, antlions), Megaloptera (alderflies, dobsonflies), and Raphidioptera (snakeflies), and includes ...

Holometabola