''Uria'' is a genus of

seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...

s in the

auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...

family known in

Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...

guillemot Guillemot is the common name for several species of seabird in the Alcidae or auk family (part of the order Charadriiformes). In British use, the term comprises two genera: ''Uria'' and ''Cepphus''. In North America the ''Uria'' species are ...

s, in most of

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

as murres, and in

Newfoundland and Labrador Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...

as turr. These are medium-sized birds with mainly brown or black plumage in the breeding season. They breed on the coasts of the northern

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

and

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...

oceans.

Taxonomy

The

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

''Uria'' was introduced by the French zoologist

Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...

in 1760 with the

common murre The common murre or common guillemot (''Uria aalge'') is a large auk. It has a circumpolar distribution, occurring in low-Arctic and boreal waters in the North Atlantic and North Pacific. It spends most of its time at sea, only coming to land t ...

(''Uria aalge'') as the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

. The genus name is from

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...

''ouria'', a waterbird mentioned by

Athenaeus Athenaeus of Naucratis (; grc, Ἀθήναιος ὁ Nαυκρατίτης or Nαυκράτιος, ''Athēnaios Naukratitēs'' or ''Naukratios''; la, Athenaeus Naucratita) was a Greek rhetorician and grammarian, flourishing about the end of ...

. The English "guillemot" is from French ''guillemot'' probably derived from ''Guillaume'', "William". "Murre" is of uncertain origins, but may imitate the call of the common guillemot. ''Uria'' auks are relatives of the

razorbill The razorbill, razor-billed auk, or lesser auk (''Alca torda'') is a colonial seabird and the only extant member of the genus '' Alca'' of the family Alcidae, the auks. It is the closest living relative of the extinct great auk (''Pinguinis im ...

little auk The little auk or dovekie (''Alle alle'') is a small auk, the only member of the genus ''Alle''. ''Alle'' is the Sami name of the long-tailed duck; it is onomatopoeic and imitates the call of the drake duck. Linnaeus was not particularly f ...

and the

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

great auk The great auk (''Pinguinus impennis'') is a species of flightless alcid that became extinct in the mid-19th century. It was the only modern species in the genus ''Pinguinus''. It is not closely related to the birds now known as penguins, wh ...

and together make up the

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...

Alcini. Despite the similar British common names, they are not so closely related to the '' Cepphus'' guillemots, which form the tribe Cepphini.

Extant species

The genus contains two species:

Description

These birds breed in large

colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...

on coastal cliffs, laying single elongated conical eggs directly on cliff ledges. They

move Move may refer to: People * Daniil Move (born 1985), a Russian auto racing driver Brands and enterprises * Move (company), an online real estate company * Move (electronics store), a defunct Australian electronics retailer * Daihatsu Move Go ...

south in winter to keep in ice-free waters. They dive for food from the surface, swimming underwater and being among the deepest divers of all birds, using their stubby wings to routinely dive to more than 100 meters, and feeding on

fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...

and

crustacea Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group c ...

ns, also some

mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is est ...

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...

s and plant material. Adult birds are black or brown on the head, neck, back and wings with white underparts. The bill is long and pointed. They have a small rounded black tail. The lower face becomes white in winter. The flight is strong and direct, and these species have fast wing beats due to the short wings. ''Uria'' guillemots produces a variety of harsh cackling calls at the breeding colonies, but are silent at sea.

Fossils

Some prehistoric ''Uria'' species are also known: * ''Uria bordkorbi'' Howard, 1981 (

Monterey Monterey (; es, Monterrey; Ohlone: ) is a city located in Monterey County on the southern edge of Monterey Bay on the U.S. state of California's Central Coast. Founded on June 3, 1770, it functioned as the capital of Alta California under bot ...

or Sisquoc, Late

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

of Lompoc, USA) * ''Uria affinis'' (Marsh, 1872) (Late

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

of E USA) - possibly a subspecies of ''U. lomvia'' * ''Uria paleohesperis'' Howard, 1982 (Late Miocene of W USA) * ''Uria onoi'' Watanabe, Matsuoka and Hasegawa, 2016 (Middle-Late Pleistocene of Japan) ''U. brodkorbi'' is interesting insofar as it is the only known occurrence of the Alcini tribe in the temperate to subtropical

, except for the very fringe of the range of ''U. aalge''. It suggests that the ''Uria'' species, which are the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to all the other Alcini, and like them are usually believed to have evolved in the Atlantic, may have evolved in the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...

or possibly close to the

Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...

. The modern Pacific distribution would then be part of a later

arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...

expansion, whereas most other auk lineages form

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...

s with a continuous range in the Pacific, from arctic to subtropical waters.

References

External links

''Hunting ‘Turr’ in Newfoundland’s Frigid Waters'', New York Times, 4Dec. 201

{{Authority control Auks Atlantic auks Pacific auks Guillemots Bird genera