Mullus barbatus
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''Mullus barbatus'' (red mullet) is a species of
goatfish The goatfishes are perciform fish of the family Mullidae. The family is also sometimes referred to as the red mullets, which also refers more narrowly to the genus '' Mullus''. The family name and the English common name mullet derived from La ...
found in the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...
,
Sea of Marmara The Sea of Marmara,; grc, Προποντίς, Προποντίδα, Propontís, Propontída also known as the Marmara Sea, is an inland sea located entirely within the borders of Turkey. It connects the Black Sea to the Aegean Sea via the ...
, the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Roma ...
and the eastern
North Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe an ...
, where its range extends from
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
to
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
. They are fished, mostly by trawling, with the flesh being well regarded. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed their conservation status as being of "
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
".


Taxonomy

This fish was first described in 1758 as ''Mullus barbatus'' by the Swedish zoologist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
. FishBase currently recognizes two subspecies, though the validity of ''M. b. ponticus'' is uncertain: * ''M. b. barbatus'' (red mullet)
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
(found throughout most of its range) * ''M. b. ponticus'' (blunt-snouted mullet) Essipov, 1927 (found in the Black Sea and the
Sea of Azov The Sea of Azov ( Crimean Tatar: ''Azaq deñizi''; russian: Азовское море, Azovskoye more; uk, Азовське море, Azovs'ke more) is a sea in Eastern Europe connected to the Black Sea by the narrow (about ) Strait of Kerch, ...
)


Description

The red mullet can grow to a standard length of , but a more common length is about half that. The body is somewhat laterally compressed. The snout is short and steep and there are no spines on the operculum. The upper jaw is toothless, but there are teeth on the roof of the mouth and on the lower jaw. A pair of moderately long
barbels In fish anatomy and turtle anatomy, a barbel is a slender, whiskerlike sensory organ near the mouth. Fish that have barbels include the catfish, the carp, the goatfish, the hagfish, the sturgeon, the zebrafish, the black dragonfish and some ...
on the chin do not exceed the pectoral fins in length. The first dorsal fin has eight spines (the first one tiny) and the second dorsal fin has one spine and eight soft rays. This fish is rose-pink, without distinctive markings on its fins.


Distribution and habitat

The red mullet is found in the Mediterranean Sea, the Black Sea and the northeastern and central eastern Atlantic Ocean, where its range extends from Scandinavia southwards to Senegal, including
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
,
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
and
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
. It is a
demersal The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...
fish and occurs at depths ranging from over muddy, sandy or gravel bottoms.


Ecology

The red mullet is carnivorous, the diet consisting mainly of
polychaete worms Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are m ...
,
bivalve molluscs Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
and
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s. The barbels are sensory organs and are used to help locate prey. Both young and adult mullet are preyed on by various fish including the
angler fish The anglerfish are fish of the teleost order (biology), order Lophiiformes (). They are Osteichthyes, bony fish named for their characteristic mode of predation, in which a modified luminescent Fish_fin#Ray-fins, fin ray (the esca or illicium) a ...
(''Lophius piscatorius''), the
thornback ray The thornback ray (''Raja clavata''), or thornback skate, is a species of ray fish in the family Rajidae. Distribution The Thornback ray is found in the Atlantic coastal waters of Europe and western Africa. It is also present from south Afri ...
(''Raja clavata''), the
common stingray The common stingray (''Dasyatis pastinaca'') is a species of stingray in the family Dasyatidae, found in the northeastern Atlantic Ocean and the Mediterranean and Black Seas. It typically inhabits sandy or muddy habitats in coastal waters shall ...
(''Dasyatis pastinaca''), the
school shark The school shark (''Galeorhinus galeus'') is a houndshark of the family Triakidae, and the only member of the genus ''Galeorhinus''. Common names also include tope, tope shark, snapper shark, and soupfin shark. It is found worldwide in temperate ...
(''Galeorhinus galeus''), the
John Dory John Dory, St Pierre or Peter's fish, refers to fish of the genus ''Zeus'', especially ''Zeus faber'', of widespread distribution. It is an edible demersal coastal marine fish with a laterally compressed olive-yellow body which has a large dark ...
(''Zeus faber'') and the
European hake ''Merluccius merluccius'', the European hake, is a merluccid hake of the genus ''Merluccius''. Other vernacular names include Cornish salmon and herring hake. It is a predatory species which was often netted alongside one of its favoured prey, th ...
(''Merluccius merluccius''). Breeding takes place in the spring and summer, with spawning occurring in April and May in the Adriatic Sea, at depths between . The larvae soon move to shallower depths and are
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
, as are the juveniles at first. At a length of about the juveniles move to the coast and become demersal, often congregating in estuaries, and sometimes swimming a short distance upstream. Later they disperse to muddy, sandy or gravelly substrates, becoming sexually mature at a length of during their first year of life.


Status

The flesh of the red mullet is much esteemed and it is the target of fisheries, especially in the Mediterranean Sea, Black Sea and northeastern and central eastern Atlantic Ocean. It is mainly caught by trawling, but also with trammel nets,
gill net Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...
s and hook and line, and in artisanal fisheries with traps and spears. In the Mediterranean there are signs of overfishing, and many of the fish caught are shorter than in total length, and being under two years old, are not yet sexually mature. For conservation of the species in the Mediterranean, the breeding grounds and nursery areas need to be protected. It is also heavily fished off the coast of northwestern Africa. Here the fish are caught by local artisan fishermen as well as by foreign industrial fleets; they may be the targeted species or may be
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
in
hake The term hake refers to fish in the: * Family Merlucciidae of northern and southern oceans * Family Phycidae (sometimes considered the subfamily Phycinae in the family Gadidae) of the northern oceans Hake Hake is in the same taxonomic order (Gad ...
,
cephalopod A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ...
or
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
fisheries, but the catch statistics are not subdivided by species. There is also thought to be overfishing in the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Roma ...
. The fish is currently listed by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
as being of "
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
" because it has a wide range, occurs at depths down to and is expanding its range northwards as a result of rising sea temperatures.


References

{{Taxonbar, from=Q7171
red mullet Red is the color at the long wavelength end of the visible spectrum of light, next to orange and opposite violet. It has a dominant wavelength of approximately 625–740 nanometres. It is a primary color in the RGB color model and a seconda ...
Fish of the Atlantic Ocean Fish of the Black Sea Fish of the Mediterranean Sea Fish of Europe Fauna of Macaronesia Marine fauna of North Africa Fish described in 1758 Taxa named by Carl Linnaeus