''Mollesaurus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of large ophthalmosaurine

ophthalmosaurid Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the ...

ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, alt ...

known from northwestern

Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...

Etymology

''Mollesaurus'' was named by Marta S. Fernández in

1999 File:1999 Events Collage.png, From left, clockwise: The funeral procession of King Hussein of Jordan in Amman; the 1999 İzmit earthquake kills over 17,000 people in Turkey; the Columbine High School massacre, one of the first major school shoot ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

is ''Mollesaurus periallus''. The generic name is derived from the name of the

Los Molles Formation The Los Molles Formation is a geologic formation of Early to Middle Jurassic age, located at northern and central part of Neuquén Basin at Mendoza Shelf in Argentina. It is overlain by the Niyeu–Lajas Formation.McIlroy et al., 2005 Descrip ...

, where the holotype was collected, and ''sauros'',

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for "lizard". The specific name is derived from ''periallos'',

for "before all others", in reference to the fact that it is the oldest ophthalmosaurid and one of the oldest thunnosaurs.

History of study

''Mollesaurus'' is known from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

MOZ 2282 V, articulated partial

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

which preserved partial

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...

and most of the

vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...

. It was collected in the Chacaico Sur locality from the ''Emileia giebeli'' ammonoid zone of the

, Cuyo Group, dating to the early

Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stra ...

stage of the

Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively ...

, about 171.6-170

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago ...

. ''Mollesaurus'', along with '' Chacaicosaurus cayi'' which was found at the same locality, are the only diagnostic ichthyosaur specimens from the

Aalenian The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.1 Ma to about 170.3 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian. Stratigraphi ...

Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. ...

interval. It was found near Zapala city of the Neuquén Basin. Maisch and Matzke (2000) regarded ''Mollesaurus'' to be a species of ''

Ophthalmosaurus ''Ophthalmosaurus'' (meaning "eye lizard" in Greek) is an ichthyosaur of the Jurassic period (165–150 million years ago). Possible remains from the Cretaceous, around 145 million years ago, are also known. It was a relatively medium-sized ich ...

''. However, all recent

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analyses found that ''Mollesaurus'' is a valid genus of ophthalmosaurid. Patrick S. Druckenmiller and Erin E. Maxwell (2010) found it to be the basalmost member of the ophthalmosaurid lineage that included '' Brachypterygius'', '' Caypullisaurus'' and ''

Platypterygius ''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus ...

'' (but not ''Ophthalmosaurus''). Valentin Fischer, Michael W. Maisch, Darren Naish, Ralf Kosma, Jeff Liston, Ulrich Joger, Fritz J. Krüger, Judith Pardo Pérez, Jessica Tainsh and Robert M. Appleby (2012) found it to be the basalmost member of

Ophthalmosaurinae Ophthalmosaurinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the Middle Jurassic to the late Early Cretaceous (Bajocian - Albian) of Europe, North America and South America. Currently, the oldest and the basal ...

Phylogeny

The following cladogram shows a possible phylogenetic position of ''Mollesaurus'' in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).

References

{{Taxonbar, from=Q6896406 Ophthalmosaurinae Middle Jurassic ichthyosaurs Bajocian life Middle Jurassic reptiles of South America Jurassic Argentina Fossils of Argentina Neuquén Basin Fossil taxa described in 1999 Ichthyosauromorph genera

Etymology

History of study

Phylogeny

See also

References