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Calobatinae Calycopteryginae Eurybatinae Micropezinae Taeniapterinae

Diversity

54 genera, 5 subfamilies

Mating pair

The Micropezidae
Micropezidae
are a moderate-sized family of acalyptrate muscoid flies in the insect order Diptera, comprising about 500 species in about 50 genera and five subfamilies worldwide, (except New Zealand and Macquarie Island).[1] They are most diverse in tropical and subtropical habitats, especially in the Neotropical Region. Insects in this family are commonly called stilt-legged flies, after their characteristically long legs. The fore legs are markedly smaller than the other pairs. Mostly, they are long-bodied, often black flies, usually with infuscated (darkened) wings. Wings are reduced in the genera Calycopteryx and entirely absent in the ant-like Badisis ambulans.[1]

Contents

1 Description 2 Biology 3 Identification 4 Conservation 5 Fossil record 6 Genera 7 References 8 Further reading 9 External links

Description[edit] For terms see Morphology of Diptera Very slender, small to large (3–16 mm) flies, they have long, thin legs and narrow wings. The head is small and elongated or rounded. The antennae are small and the arista is bare or pubescent. Ocelli are present, but ocellar bristles are absent. Vibrissae are absent and the postvertical bristles are divergent or absent. Up to three pairs of frontal bristles curve forward or backward. Interfrontal bristles are absent. The wings are clear or have a smoky pattern. The costa is without interruptions. The subcosta is complete, its ending in the costa close to vein R1. The posterior basal wing cell and discoidal wing cell are sometimes fused. Crossvein BM-Cu present or (Micropezinae) absent. The abdomen is long and narrow. The tibiae lack a dorsal preapical bristle. Biology[edit]

Play media

Micropeza
Micropeza
corrigiolata

Play media

Neria cibaria

Some species, much the same as in the strongylophthalmyiid genus, Strongylophthalmyia, mimic ants; others mimic wasps and are especially similar in appearance to some ichneumonid wasps. Species of the genus Anaeropsis have stalked eyes. Little is known of the larval habits, but they are probably phytophagous or saprophagous in decayed vegetation, old manure, or fungi . Larvae of certain Mimegralla species have been found to live in the roots of ginger and other plants, under the bark of dead trees, or in other decaying material. Species of Micropeza
Micropeza
have phytophagous larvae feeding in the root nodules of leguminous plants in open habitats. Species of Rainieria
Rainieria
develop in rotting wood and are found in old forests. Adults are either predatory on small insects (for example Calobata in Britain[1]) or are attracted to excrement or decaying fruit. Adults are found on low herbage, flowers, leaves, rotting fruit, and excrement. Many species (for example those of genus Mimegralla) are known for their habit of standing motionless while waving their prominently marked front legs in front of their heads, a behavior which contributes to their mimicry of wasps. At least one species of Metopochetus (M. curvus) was observed to wave its hind legs instead, though these are not conspicuously colored.[1] Identification[edit]

Czerny. 1930. Tylidae. In: Lindner, E. (Ed.). Die Fliegen der palaearktischen Region 5, 42a, 13-16. Keys to Palaearctic species but now needs revision (in German). Séguy, E. (1934) Diptères: Brachycères. II. Muscidae
Muscidae
acalypterae, Scatophagidae. Paris: Éditions Faune de France 28. virtuelle numérique Shtakel'berg, A.A. Family Micropezidae
Micropezidae
(Tylidae in part) in Bei-Bienko, G. Ya, 1988 Keys to the insects of the European Part of the USSR Volume 5 (Diptera) Part 2 English edition. Keys to Palaearctic species but now needs revision.

Conservation[edit] Calycopteryx mosleyi, found on the Kerguelen Islands
Kerguelen Islands
and Heard Island, is associated with the Kerguélen cabbage (Pringlea antiscorbutica, Brassicaceae). As this plant is being destroyed by introduced rabbits, the fly is considered vulnerable.[1] The larvae of Badisis ambulans
Badisis ambulans
live in the pitchers of the endangered Albany pitcher plant (Cephalotus follicularis, Cephalotaceae).[1] Fossil record[edit] Several fossil species have been found in Baltic amber, probably from the Late Eocene
Late Eocene
(about 36 million years old). However, most were washed onto beaches after wearing from Tertiary strata, making their age uncertain. Two fossil genera are presently recognized, Cypselosomatites Hennig, 1965 and the advanced Electrobata Hennig, 1965 which may be more than one genus however. Genera[edit] This list is compiled from the BioSystematic Database of World Diptera and probably complete as of January 2007.

Anaeropsis Bigot, 1866 Badisis
Badisis
McAlpine, 1990 Calobata Meigen, 1803 Calobatella Mik, 1898 Calobatina Enderlein, 1922 Calosphen Hennig, 1934 Calycopteryx Eaton, 1875 Cardiacephala Macquart, 1843 Cephalosphen Hennig, 1934 Chaetotylus Hendel, 1932 Cliobata Enderlein, 1923 Cnodacophora Czerny, 1930 Compsobata Czerny, 1930 Cothornobata Czerny, 1932 Courtoisia Barraclough, 1993 Crepidochetus Enderlein, 1922 Crosa Steyskal, 1952 Cryogonus Cresson, 1926 Ectemnodera Enderlein, 1922 Electrobata Hennig, 1965 Erythromyiella Hennig, 1935 Eurybata Osten Sacken, 1882 Globomyia Hennig, 1935 Globopeza Marshall, 2005 Glyphodera Enderlein, 1922 Grallipeza
Grallipeza
Rondani, 1850 Grammicomyia Bigot, 1859 Hemichaeta Hennig, 1934 Hoplocheiloma Cresson, 1926 Mesoconius Enderlein, 1922 Metasphen Frey, 1927 Metopochetus Enderlein, 1922 Micropeza
Micropeza
Meigen, 1803 Mimegralla Rondani, 1850 Mimomyrmecia Frey, 1927 Neograllomyia Hendel, 1933 Neria Robineau-Desvoidy, 1830 Nestima Osten Sacken, 1881 Notenthes Marshall, 2002 Papeza McAlpine, 1975 Paramimegralla Hennig, 1937 Parasphen Enderlein, 1922 Plocoscelus Enderlein, 1922 Poecilotylus Hennig, 1934 Pseudeurybata Hennig, 1934 Ptilosphen Enderlein, 1922 Rainieria
Rainieria
Rondani, 1843 Scipopus Enderlein, 1922 Steyskalia Aczel, 1959 Stiltissima Barraclough, 1991 Taeniaptera
Taeniaptera
Macquart, 1835 Tenthes Cresson, 1930 Trepidarioides Frey, 1927 Zelatractodes Enderlein, 1922

References[edit]

^ a b c d e f McAlpine, D.K. (1998). Review of the Australian stilt flies (Diptera: Micropezidae) with a phylogenetic analysis of the family. Invertebrate Taxonomy 12:55–134. doi:10.1071/IT96018 (with key to Australian species)

Further reading[edit]

This article includes a list of references, but its sources remain unclear because it has insufficient inline citations. Please help to improve this article by introducing more precise citations. (April 2009) (Learn how and when to remove this template message)

Andersson, H., 1989 Taxonomic notes on the Fennoscandian Micropezidae.Notulae Entomologicae, 69:153–162. Brown, B. V., A. Borkent, J.M. Cumming, D.M. Wood, N.E. Woodley, and M. Zumbado. 2010. Manual of Central American Diptera, Volume 2. NRC Research Press. Hennig, W. 1934. Revision der Tyliden (Dipt., Acalypt.). I. Teil: die Taeniapterinae Amerikas [part]. Stett. Entomol. Ztg. 95: 65–108, 294–330. [Publication split: . 65–108 ; . 294–330 . Concludes in Hennig, 1935b.] Hennig, W. 1935a. Revision der Tyliden (Dipt., Acalypt.). II. Teil: die ausseramerikanischen Taeniapterinae, die Trepidariinae und Tylinae. Allgemeines über die Tyliden. Zugleich ein Beitrag zu den Ergebnissen der Sunda-expedition Rensch, 1927 [part]. Konowia 14: 68–92. Hennig, W. 1935b. Revision der Tyliden (Dipt., Acalypt.). I. Teil: die Taeniapterinae Amerikas [concl.]. Stett. Entomol. Ztg. 96: 27–67. Hennig, W. 1935c. Revision der Tyliden (Dipt., Acalypt.). II. Teil: die ausseramerikanischen Taeniapterinae, die Trepidariinae und Tylinae. Allgemeines über die Tyliden. Zugleich ein Beitrag zu den Ergebnissen der Sunda-expedition Rensch, 1927 [part]. Konowia 14: 192–216, 289–310. [Publication split: . 192–216 ; 289–310. Concludes in Hennig, 1936.] Hennig, W. 1936. Revision der Tyliden (Dipt., Acalypt.). II. Teil: die ausseramerikanischen Taeniapterinae, die Trepidariinae und Tylinae. Allgemeines über die Tyliden. Zugleich ein Beitrag zu den Ergebnissen der Sunda-expedition Rensch, 1927 [concl.]. Konowia 15: 129–44, 201–39. [Publication split: . 129–44 ; . 201–39. ] McAlpine, D.K., 1975. The subfamily classification of the Micropezidae and the genera of Eurybatinae (Diptera: Schizophora). Journal of Entomology (B) 43: 231–245. Steyskal, G.C. 1952b. Australasian stilt-legged flies (Diptera: Tylidae) in the United States National Museum. Proc. U.S. Natl. Mus. 102 [= No. 3294): 161–80. (26 February) Steyskal, G.C. 1964. Larvae of Micropezidae
Micropezidae
(Diptera), including two species that bore in ginger roots. Ann. Entomol. Soc. Am. 57: 292–96. (15 May) Steyskal, G.C. 1977a. Family Micropezidae
Micropezidae
(Tylidae), . 12–20. In: Delfinado, M.D. & D.E. Hardy, eds., A catalog of the Diptera
Diptera
of the Oriental Region. Volume III. Suborder Cyclorrhapha (excluding Division Aschiza). University Press of Hawaii, Honolulu. x + 854 p. Verbeke, J. 1951. Taenapterinae (Diptera:Cyclorrhapha) Fam. Micropezidae. Explor. Parc. natn. Albert Miss. G.F. de Witte 72:1–106.

External links[edit]

Wikimedia Commons has media related to Micropezidae.

Wikispecies
Wikispecies
has information related to Micropezidae

Family Micropezidae
Micropezidae
at EOL Stilt-legged fly diagnostic photographs Key to U.S.A. Micropezidae European (including Russia) species list Nearctic (North America) species list BugGuide Images at BugGuide Images at Diptera.info

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Extant Diptera
Diptera
families

Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera Superorder: Endopterygota

Suborder Nematocera

Axymyiomorpha

Axymyiidae

Culicomorpha

Culicoidea

Dixidae
Dixidae
(meniscus midges) Corethrellidae
Corethrellidae
(frog-biting midges) Chaoboridae
Chaoboridae
(phantom midges) Culicidae (mosquitoes)

Chironomoidea

Thaumaleidae
Thaumaleidae
(solitary midges) Simuliidae (black flies) Ceratopogonidae
Ceratopogonidae
(biting midges) Chironomidae
Chironomidae
(non-biting midges)

Blephariceromorpha

Blephariceridae
Blephariceridae
(net-winged midges) Deuterophlebiidae (mountain midges) Nymphomyiidae

Bibionomorpha

Bibionoidea

Bibionidae
Bibionidae
(march flies, lovebugs)

Anisopodoidea

Anisopodidae
Anisopodidae
(wood gnats)

Sciaroidea (fungus gnats)

Bolitophilidae Diadocidiidae Ditomyiidae Keroplatidae Mycetophilidae Sciaridae
Sciaridae
(dark-winged fungus gnats) Cecidomyiidae
Cecidomyiidae
(gall midges)

Psychodomorpha

Scatopsoidea

Canthyloscelidae Perissommatidae Scatopsidae
Scatopsidae
(minute black scavenger flies, or dung midges)

Psychodoidea

Psychodidae (moth flies)

Ptychopteromorpha

Ptychopteridae
Ptychopteridae
(phantom crane flies) Tanyderidae (primitive crane flies)

Tipulomorpha

Trichoceroidea

Trichoceridae
Trichoceridae
(winter crane flies)

Tipuloidea

Pediciidae
Pediciidae
(hairy-eyed craneflies) Tipulidae (crane flies)

Suborder Brachycera

Asilomorpha

Asiloidea

Apioceridae (flower-loving flies) Apsilocephalidae Apystomyiidae Asilidae
Asilidae
(robber flies) Bombyliidae
Bombyliidae
(bee flies) Evocoidae Hilarimorphidae (hilarimorphid flies) Mydidae (mydas flies) Mythicomyiidae Scenopinidae
Scenopinidae
(window flies) Therevidae
Therevidae
(stiletto flies)

Empidoidea

Atelestidae Hybotidae
Hybotidae
(dance flies) Dolichopodidae
Dolichopodidae
(long-legged flies) Empididae
Empididae
(dagger flies, balloon flies)

Nemestrinoidea

Acroceridae
Acroceridae
(small-headed flies) Nemestrinidae
Nemestrinidae
(tangle-veined flies)

Muscomorpha

Aschiza

Platypezoidea

Phoridae
Phoridae
(scuttle flies, coffin flies, humpbacked flies) Opetiidae
Opetiidae
(flat-footed flies) Ironomyiidae (ironic flies) Lonchopteridae
Lonchopteridae
(spear-winged flies) Platypezidae
Platypezidae
(flat-footed flies)

Syrphoidea

Syrphidae (hoverflies) Pipunculidae
Pipunculidae
(big-headed flies)

Schizophora

Acalyptratae

Conopoidea

Conopidae
Conopidae
(thick-headed flies)

Tephritoidea

Pallopteridae
Pallopteridae
(flutter flies) Piophilidae
Piophilidae
(cheese flies) Platystomatidae
Platystomatidae
(signal flies) Pyrgotidae Richardiidae Tephritidae
Tephritidae
(peacock flies) Ulidiidae
Ulidiidae
(picture-winged flies)

Nerioidea

Cypselosomatidae Micropezidae
Micropezidae
(stilt-legged flies) Neriidae
Neriidae
(cactus flies, banana stalk flies)

Diopsoidea

Diopsidae (stalk-eyed flies) Gobryidae Megamerinidae Nothybidae Psilidae
Psilidae
(rust flies) Somatiidae Strongylophthalmyiidae Syringogastridae Tanypezidae

Sciomyzoidea

Coelopidae
Coelopidae
(kelp flies) Dryomyzidae Helosciomyzidae Ropalomeridae Huttoninidae Heterocheilidae Phaeomyiidae Sepsidae
Sepsidae
(black scavenger flies) Sciomyzidae
Sciomyzidae
(marsh flies)

Sphaeroceroidea

Chyromyidae Heleomyzidae Sphaeroceridae
Sphaeroceridae
(small dung flies) Nannodastiidae

Lauxanioidea

Celyphidae
Celyphidae
(beetle-backed flies) Chamaemyiidae
Chamaemyiidae
(aphid flies) Lauxaniidae

Opomyzoidea

Agromyzidae
Agromyzidae
(leaf miner flies) Anthomyzidae Asteiidae Aulacigastridae (sap flies) Clusiidae
Clusiidae
(lekking, or druid flies) Fergusoninidae Marginidae Neminidae Neurochaetidae (upside-down flies) Odiniidae Opomyzidae Periscelididae Teratomyzidae Xenasteiidae

Ephydroidea

Camillidae Curtonotidae
Curtonotidae
(quasimodo flies) Diastatidae
Diastatidae
(bog flies) Ephydridae
Ephydridae
(shore flies) Drosophilidae
Drosophilidae
(vinegar and fruit flies)

Carnoidea

Acartophthalmidae Australimyzidae Braulidae
Braulidae
(bee lice) Canacidae
Canacidae
(beach flies) Carnidae Chloropidae
Chloropidae
(frit flies) Cryptochaetidae Inbiomyiidae Milichiidae
Milichiidae
(freeloader flies)

Lonchaeoidea

Cryptochetidae Lonchaeidae
Lonchaeidae
(lance flies)

Calyptratae

Muscoidea

Anthomyiidae
Anthomyiidae
(cabbage flies) Fanniidae
Fanniidae
(little house flies) Muscidae
Muscidae
(house flies, stable flies) Scathophagidae
Scathophagidae
(dung flies)

Oestroidea

Calliphoridae
Calliphoridae
(blow-flies: bluebottles, greenbottles) Mystacinobiidae ( New Zealand
New Zealand
batfly) Oestridae (botflies) Rhinophoridae Sarcophagidae (flesh flies) Tachinidae
Tachinidae
(tachina flies)

Hippoboscoidea

Glossinidae (tsetse flies) Hippoboscidae
Hippoboscidae
(louse flies) Mormotomyiidae
Mormotomyiidae
(frightful hairy fly) Nycteribiidae
Nycteribiidae
(bat flies) Streblidae
Streblidae
(bat flies)

Stratiomyomorpha

Stratiomyoidea

Pantophthalmidae
Pantophthalmidae
(timber flies) Stratiomyidae
Stratiomyidae
(soldier flies) Xylomyidae
Xylomyidae
(wood soldier flies)

Tabanomorpha

Rhagionoidea

Austroleptidae Bolbomyiidae Rhagionidae
Rhagionidae
(snipe flies)

Tabanoidea

Athericidae
Athericidae
(water snipe flies) Oreoleptidae Pelecorhynchidae Tabanidae (horse and deer flies)

Vermileonomorpha

Vermileonoidea

Vermileonidae

Xylophagomorpha

Xylophagoidea

Xylophagidae
Xylophagidae
(awl flies)

List of families of Diptera

v t e

Micropezidae

Genera

Anaeropsis Badisis Calobata Calobatella Calobatina Calosphen Calycopteryx Cardiacephala Cephalosphen Chaetotylus Cliobata Cnodacophora Compsobata Cothornobata Courtoisia Crepidochetus Crosa Cryogonus Ectemnodera Electrobata Erythromyiella Eurybata Globomyia Globopeza Glyphodera Grallipeza Grammicomyia Hemichaeta Hoplocheiloma Mesoconius Metasphen Metopochetus Micropeza Mimegralla Mimomyrmecia Neograllomyia Neria Nestima Notenthes Papeza Paramimegralla Parasphen Plocoscelus Poecilotylus Pseudeurybata Ptilosphen Rainieria Scipopus Steyskalia Stiltissima Taeniaptera Tenthes Trepidarioides Zelatractodes

Taxon identifiers

Wd: Q1782917 BugGuide: 134 EoL: 431 EPPO: 1MICPF Fauna Europaea: 10924 Fossilworks: 139219 GBIF: 9495 iNaturalist: 132187 ITIS: 142035 NCBI: 11

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